www.geologicacarpathica.sk
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
, AUGUST 2011, 62, 4, 333—343 doi: 10.2478/v10096-011-0025-8
Introduction
The identification of the Cretaceous-Paleogene (K-T) bound-
ary in turbiditic deposits is problematic, mostly because of
the absence of index fossils, especially planktonic foramini-
fers, and because of redeposition processes disturbing the
stratigraphic record. Only a few attempts can be found in the
literature, where the boundary is determined within some
portion of the section (Melinte 1999; Bubík et al. 2002;
Chira et al. 2009). Gasiński & Uchman (2009) put the
boundary within a 3 m thick interval of the Ropianka Forma-
tion (Upper Cretaceous—Paleocene) section in the Skole
Nappe in the Polish Flysch Carpathians. The boundary was
identified above the last occurrence of the planktonic Creta-
ceous foraminifers, including the index taxon Abathom-
phalus mayaroensis, and below the occurrence of benthic
foraminifers typical of the Paleogene. As a result of new in-
vestigations in a better exposed section of the Ropianka For-
mation in the adjacent thrust sheet, not only the Cretaceous
but also the Paleogene index planktonic foraminiferids have
been found that enabled us to recognize the K-T boundary
with the accuracy to two beds for the first time in these tur-
biditic deposits. Micropaleontological analysis of this sec-
tion, and the K-T boundary in particular, are the main aim of
this paper.
Geological setting
The Skole Nappe is the most external major nappe of the
Polish Flysch Carpathians (Fig. 1A). It is composed of Low-
er Cretaceous—Miocene deep-sea, mostly flysch sediments
The Cretaceous-Paleogene boundary in turbiditic deposits
identified to the bed: a case study from the Skole Nappe
(Outer Carpathians, southern Poland)
M. ADAM GASIŃSKI and ALFRED UCHMAN
Institute of Geological Sciences, Jagiellonian University, Oleandry Str. 2a, 30-063 Kraków, Poland;
adam.gasinski@uj.edu.pl; alfred.uchman@uj.edu.pl
(Manuscript received November 3, 2010; accepted in revised form February 4, 2011)
Abstract: The Cretaceous-Paleogene (K-T) boundary has been recognized in turbiditic sediments of the Ropianka
Formation in the Skole Nappe (Bąkowiec section) on the basis of planktonic foraminiferids with an accuracy of 40 cm.
Such precise determination of the K-T boundary for the first time in the Carpathians and in turbiditic flysch sediments
in general was possible due to the successive occurrence of the Early Paleocene planktonic taxa of the P1 Zone above the
latest Maastrichtian Abathomphalus mayaroensis Zone with the Racemiguembelina fructicosa Subzone. The trends in
composition of the latest Maastrichtian foraminiferal assemblages are similar to the Gaj section from the adjacent thrust
sheet, probably due to the influence of the same paleoenvironmental factors.
Key words: Paleocene, Maastrichtian, K-T boundary, Carpathians, paleoecology, biostratigraphy, turbidites,
foraminiferids.
that accumulated in the Skole Basin, a segment of the north-
ern Neotethys, and were folded and thrust northward during
the Miocene. The Upper Cretaceous—Paleocene sediments
are distinguished as the Ropianka Formation, which was also
named the Inoceramian Beds for a long time (Kotlarczyk
1978). They are overlain by the Eocene Variegated Shale
Formation (Rajchel 1990). For the complex history of re-
search on the Ropianka Formation in the Skole Nappe see
Kotlarczyk (1978). SE of Rzeszów, Wdowiarz (1949) distin-
guished the lower, middle and upper levels in the Ropianka
Formation (his Inoceramian Beds), which are altogether
500 m thick. Kotlarczyk (1978) subdivided the Ropianka
Formation into the Cisowa Member (Turonian—Lower Cam-
panian), Wiar Member (Lower Campanian—Lower Maas-
trichtian), Leszczyny Member (Lower Maastrichtian—Lower
Paleocene) and Wola Korzeniecka Member (Paleocene). For
further details see Gasiński & Uchman (2009) and references
therein.
The lower part of the studied section (GPS coordinates:
N49°59.040
’; E022°14.910’; ±9 m) starts along an un-
named stream, a tributary of the Handzlowski Potok Stream
(Sawa River in the lower part), the gorge of which is incised
into the southern slope of the Patria Hill (426 m a.s.l.) along
the border between the Husów and Handzlówka villages,
and continues along the Handzlowski Potok Stream in the
Bąkowiec forest on the territory of the Husów village
(Fig. 1B). The outcrops are small and isolated (Figs. 1B, 2),
but consequent strikes and dips of beds suggest a monoclinal
structure forming a part of the Husów Thrust Sheet. The
thrust sheet is disturbed by Variegated Shale occurring in a
narrow stripe stretching along the Handzlowski Potok
Stream (Wdowiarz 1949), which suggests an internal thrust.
334
GASIŃSKI and UCHMAN
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 4, 333—343
Some tectonic reductions are possible here.
Nevertheless, the outcrops have stratigraphic
continuity.
The lower part of the section is composed
of turbiditic medium, rarely thick or thin
beds, which contain calcareous sandstones at
the base and thick layers of grey and grey-
bluish marls and marly siltstones to mud-
stones at the top (Fig. 2). In the middle part
of the section, about 4 m thick debris flow
deposits containing different marls, dark
shale and sandstone clasts occur. Higher up,
the contribution of thick sandstone and mar-
ly beds increases. Along the Handzlowski
Potok Stream, where the upper part of the
section is exposed, thin and medium beds
prevail and marly layers are replaced by cal-
careous shales in the higher part of this inter-
val. The K-T boundary was identified here
in a natural scarp on the right side of the
stream, in a 15 m thick interval, which is not
tectonically disturbed (GPS coordinates:
N49°58.706
’; E022°14.879’; ±11 m). Iso-
lated outcrops of red, non-calcareous shales,
with intercalations of rare sandstones and
green shales along the stream belong already
to the Variegated Shale Formation. Down
the stream, massive or indistinctly bedded
calcareous mudstones occur. They probably
belong to the Babica Clay (Paleocene),
which is a unit within the Variegated Shale
Formation, composed mostly of mud flow
deposits (Kotlarczyk 1978; Rajchel 1990
and references therein).
The boundary interval (Fig. 3) includes
two turbiditic-hemipelagic rhythms. The
first rhythm is composed of a 15 cm-thick,
fine-grained, grey sandstone with rusty co-
louration, which rests on a grey marlstone,
displays a sharp base and a transition to
greenish-grey marly mudstone at the top.
The marly mudstone is 8.5 cm thick and dis-
plays rusty spots. The second rhythm con-
tains very fine-grained, sharply based grey
sandstone, 5.5—6 cm thick, which passes
into a greenish-grey marlstone. The marl-
stone is 8—11 cm thick. It is overlain by a
loaded, 3.5—7.5 cm thick, very fine-grained,
laminated, grey sandstone passing into a
greenish-grey mudstone.
Stratigraphically, the studied deposits are
an equivalent of the Leszczyny Member and,
in its lowest part, possibly of the uppermost
part of the Wiar Member of the Ropianka
Formation sensu Kotlarczyk (1978, 1988).
The lithology of the Leszczyny Member
is very variable and includes marl-rich
olistostromes. It is not clear whether the
Wiar Member or the Leszczyny Member can
Fig. 1. Location map. A – Location of the study area in the Skole Nappe (simplified
after Kotlarczyk 1988). B – Map of the study area. Tectonic unit designation and
range of the Variegated Shale after Wdowiarz (1949), completed and modified. Loca-
tion of samples (Bak 1, etc.) and orientation of beds indicated.
335
THE CRETACEOUS-PALEOGENE BOUNDARY IN TURBIDITIC DEPOSITS (OUTER CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 4, 333—343
Fig. 2. Lithological columns with location of the samples.
336
GASIŃSKI and UCHMAN
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 4, 333—343
be distinguished in the study area, however a general fining
up section is one of the typical features of the Leszczyny
Member (Kotlarczyk 1978) and these members are recog-
nized west of the study area (J. Kotlarczyk, personal commu-
nication, 2008).
The K-T boundary in turbiditic sediments of the
Carpathians
Globally, the Cretaceous-Paleocene (K-T) boundary is pre-
cisely located at the base of the so-called “boundary level”,
as defined mainly in pelagic and hemipelagic sequences
(MacLeod & Keller 1996; Apellániz at al. 1997; Kaiho &
Lamolda 1999; Arenillas et al. 2004; Keller 2004; Molina et
al. 2006, and others). This boundary is situated between the
Abathomphalus mayaroensis (latest Maastrichtian) and Guem-
belitria cretacea (earliest Paleocene) Biozones (Robaszynski
& Caron 1995; Premoli-Silva et al. 2004, and others).
In turbiditic dominated sequences of the deep Alpine
flysch basins, the K-T boundary is very difficult to identify
due to the rare occurrence of index planktonic foraminiferids
and strong redeposition causing the occurrence of mixed for-
aminiferal assemblages. Only in a few cases, the identifica-
tion was narrowed to some relatively thin intervals. The
boundary occurs within about a meter thick interval in the
Magura Unit in Moravia, Czech Republic, based on dinocyst
assemblages (Bubík et al. 2002). In the Romanian Car-
pathians, it was identified within tens of meters by means of
the calcareous nannoplankton and foraminiferids (Melinte
1999; Chira et al. 2009).
The K-T boundary in the studied section
Recent studies of the Ropianka Formation of the Skole Unit
in the Husów area (Gaj section), Polish Carpathians (Gasiński
& Uchman 2009), based on 25 samples, allowed recognition
of the latest Maastrichtian Abathomphalus mayaroensis stan-
dard Biozone and overlying Paleogene deposits. The K-T
boundary has been narrowed to a 3 m thick interval, where the
latest Cretaceous planktonic foraminiferids disappear and Pa-
leogene benthic agglutinated foraminiferids (relatively long
ranging species) appear. So far, it was the most precisely iden-
tified K-T boundary in the Polish Carpathians. More precise
work in this section is limited due to its poor exposition.
In the better exposed Bąkowiec section, in the adjacent
thrust sheet, 58 samples from the fine-grained parts of tur-
biditic-hemipelagic beds were analysed in our study. The
majority of them contain relatively rich and well-preserved
foraminiferal assemblages. They can also be seen in thin sec-
tions from the sandstone beds. Similarly to the Gaj section
(Gasiński & Uchman 2009), the Gansserina gansseri and
Abathomphalus mayaroenesis standard Biozones were rec-
ognized (Fig. 4). The intermediate Racemiguembelina fructi-
cosa Zone is also distinguished, the meaning of which is
discussed below.
Moreover, the appearance of the Paleocene foraminiferids
Subbotina cancellata Blow (Fig. 5M), Subbotina triangu-
laris (White) (Fig. 5N), Eoglobigerina cf. edita (Subbotina)
(Fig. 5O) point to the earliest Paleocene zone (P1 Zone sensu
Olsson et al. 1999) (Fig. 6). The latest Maastrichtian (A.
mayaroensis) and the lowest Paleocene foraminifers occur
within a 40 cm thick interval (Figs. 3, 7) limited to two
depositional turbiditic-hemipelagic rhythms. Thus, this is the
most precisely determined K-T boundary in turbiditic
sediments.
From the base of the studied section, the Gansserina gans-
seri Zone (after Robaszynski & Caron 1995; Premoli-Silva et
al. 2004) can be recognized. Planktonic foraminiferal assem-
blages contain the index species (Fig. 4). The first appearance
(FO) of Abathomphalus mayaroensis (Bolli) is noted from the
sample Bak 2a (Figs. 4, 8M,N) and it delineates the base of
the A. mayaroensis Zone (Fig. 6). The FO of Racemiguembeli-
na fructicosa Egger (Fig. 4R) in sample Bak 2a and its last oc-
currence (LO) in sample Bak 7 (Fig. 4) allowed us to
recognize the Racemigeumbelina fructicosa Zone (Fig. 6),
which is used by some authors (Gradstein et al. 2004) as the
Partial Range Zone within the lower part of A. mayaroensis
Zone. The LO of A. mayaroensis in sample Bak 13 below the
FO of Subbotina cancellata Blow, S. triangularis (White) and
Eoglobigerina edita (Subbotina) in sample Bak 14A (Fig. 7)
has been interpreted as the latest Maastrichtian (A. mayaroensis
Zone). The latter three taxa indicate the P1 Zone of the Early
Paleocene (Olsson et al. 1999) (Fig. 6). It should be under-
lined that the K-T boundary is indentified within a sequence
that is only 40 cm thick (Fig. 3). The rusty layer overlain by
the dark boundary layer, known from pelagic and hemipelagic
sections are not recognized here, but this can be explain by the
turbiditic deposition.
The nearest located deep-sea K-T boundary outside the
Carpathians was identified in “turbiditic and partly hemipe-
Fig. 3. Detail of the K-T boundary section, with indication of sam-
ples and the boundary interval. The key samples in bold.
337
THE CRETACEOUS-PALEOGENE BOUNDARY IN TURBIDITIC DEPOSITS (OUTER CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 4, 333—343
Fig. 4. Species occurrence in the studied samples with indication of standard biozones. Occurrences of redeposited taxa in grey circles.
338
GASIŃSKI and UCHMAN
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 4, 333—343
Fig. 5. Benthic and planktonic foraminiferids of the Bąkowiec section. A – Saccammina placenta (Grzybowski), Bak 14B. B – Hormosi-
na velascoensis (Cushman), Bak 10Jc. C – Ammodiscus sp., Bak 10JA. D – Spiroplectammina sp. (cross-section), Bak 5. E – Spiro-
plectinella dentata (Alth), Bak 11. F—H – Rzehakina fissistomata (Grzybowski), Bak 10Jc. I – Rzehakina cf. inclusa (Grzybowski),
Bak 5. J – Arenobulimina preslii (Reuss), Bak 10H. K – Anomalinoides nobilis Brotzen, Bak 10H. L – Quadrimorphina allomorphi-
noides (Reuss), Bak 1. M – Subbotina cancellata Blow, Bak 14A. N – Subbotina triangularis (White), Bak 14A. O – Eoglobigerina cf.
edita (Subbotina), Bak 14A. P, U – Hedbergella holmdelensis Olsson; P – Bak 3A, U – Bak 11. R – Racemiguembelina fructicosa Egger,
Bak 6a. S – Heterohelix striata (Ehrenberg), Bak 6. T – Hedbergella monmouthensis (Olsson), Bak 3A.
339
THE CRETACEOUS-PALEOGENE BOUNDARY IN TURBIDITIC DEPOSITS (OUTER CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 4, 333—343
Fig. 6. Biostratigraphical ranges of the studied planktonic index taxa. Ranges of species and biozones plotted (combined) after Robaszynski
et al. (1984), Caron (1985), Robaszynski & Caron (1995), Olsson et al. (1999) and Premoli-Silva & Verga (2004). The grey area indicates
biozones recognized in the studied material.
lagic sediments” of the Gosau Group in the Eastern Alps
(Austria; Rotwandgraben section; Lahodynsky 1988; Peryt
et al. 1997); however, it is not clear whether the boundary is
in the turbiditic or hemipelagic sediments. In an other sec-
tion of the Gosau Group at Gams, the K-T boundary is rec-
ognized in hemipelagic sediments of a series composed of
siliciclastic and mixed siliciclastic-carbonate strata deposited
at the middle bathyal paleodepth (Egger et al. 2004, 2009).
Paleoecology
Qualitative analysis of the studied foraminiferal assem-
blages is presented in Fig. 9. Index and typical foraminifer-
ids are shown in Figs. 5, 8. The planktonic/benthic ratio,
relation of epipelagic to bathypelagic taxa among planktonic
foraminiferal assemblages, numbers of agglutinated in rela-
tion to calcareous foraminiferids within the benthic assem-
blages and amount of suspension feeders (tubular forms)
within the agglutinated foraminiferids assemblages were cal-
culated.
Charts showing the mentioned coefficients clearly fluctu-
ate (Fig. 9). Close to the K-T boundary foraminiferal assem-
blages of samples from Bak 11 to Bak 14A (14A1—14A7)
display rapid quantitative changes considered as an ecologi-
cal event. Before this event, planktonic taxa, which were
abundant (Bak 11), suddenly decrease in numbers (Bak 12,
12C). Similarly, epipelagic forms, abundant among plank-
tonic assemblages, form about 50 % in Bak 11 decrease to
about 25 % in Bak 13. Increasing number of epipelagic
(non-keeled taxa, r-strategists, opportunistic species) among
planktonic assemblages confirms a shallower depositional
environment (Gasiński 1997; Gasiński et al. 1999, 2001).
However, any sedimentological signal of shallowing sug-
gests redeposition of sediments from a shallower zone.
In contrast, agglutinated species, especially suspension
feeders, formerly relatively scarce among benthic assem-
blages (Bak 13) sharply increase in number (Bak 14A). Ag-
glutinated foraminiferids are more abundant in coarser
sediments of clearly turbiditic origin. In turbiditic sediments,
the suspension feeder morphological group dominates.
340
GASIŃSKI and UCHMAN
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 4, 333—343
Fig. 7. Species occurrence in the studied samples in the upper part of the K-T boundary interval in the Bąkowiec section, with indication of
standard biozones. The K-T boundary is located in the grey zone. Occurrences of redeposited taxa in grey circles.
341
THE CRETACEOUS-PALEOGENE BOUNDARY IN TURBIDITIC DEPOSITS (OUTER CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 4, 333—343
The correlation of quantitative charts of composition of
foraminiferal assemblages between the studied samples and
those collected from the Gaj section (Gasiński & Uchman
2009) points to their close similarity, especially in the part
Fig. 8. Planktonic foraminiferids of the Bąkowiec section. A, B – Rugoglobigerina rugosa (Plummer); A – Bak 6, B – Bak 7. C – Glo-
botruncana cf. arca Cushman, Bak 6. K, L, P – Globotruncana bulloides Vogler, Bak 11. D, I – Globotruncanita stuartiformis (Dalbiez);
D – Bak 2A, I – Bak 7. E – Contusotruncana contusa (Cushman), Bak 2A. F, G – Gansserina gansseri (Bolli), Bak 6. H – Glo-
botruncanita stuarti (de Lapparent), Bak 2A. J – Globotruncana aegyptiaca Nakkady, Bak 7. M, N – Abathomphalus mayaroensis (Bolli);
M – Bak 6, N – Bak 7. O – Archaeoglobigerina sp., Bak 13.
dated as the latest Maastrichtian (Gaj section) and K-T
boundary (studied section) (Fig. 9). It suggests that the simi-
lar factors influenced boundary section environment in this
part of the Skole Basin.
342
GASIŃSKI and UCHMAN
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 4, 333—343
Fig. 9. Quantitative analysis of the studied foraminiferal assemblages in the Bąkowiec and Gaj sections. The Gaj section data from Gasiński
& Uchman (2009).
343
THE CRETACEOUS-PALEOGENE BOUNDARY IN TURBIDITIC DEPOSITS (OUTER CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 4, 333—343
Drastic changes of foraminiferal assemblages have also
been observed in the K-T boundary sections in the pelagic/
hemipelagic facies (MacLeod & Keller 1996; Apellániz et al.
1997; Kaiho & Lamolda 1999; Arenillas et al. 2004; Keller
2004, and others).
Conclusions
1. The K-T boundary was identified in the turbiditic sedi-
ments with the accuracy of 40 cm for the first time.
2. The Gansserina gansseri, Abathomphalus mayaroensis
(Late Maastrichtian) and P1 (Early Paleocene) standard bio-
zones were recognized in the studied section on the basis of
planktonic foraminiferids.
3. The Racemiguembelina fructicosa Zone as the Partial
Range Zone within the lower part of A. mayaroensis Zone
has been determined for the first time in the Carpathians and
the flysch sediments.
4. Drastic qualitative and quantitative fluctuations among
the studied foraminiferal assemblages were recognized around
the K-T boundary similar to those indicated in the Gaj section
(next thrust sheet).
Acknowledgments: The researches were supported by the
Jagiellonian University (DS funds). The authors greatly ap-
preciate Eustoquio Molina (Zaragoza University), Ján Soták
(Slovak Academy of Sciences) and an anonymous reviewer
for their critical remarks. Waldemar Obcowski (Jagiellonian
University) prepared the photographic figures.
References
Apellániz E., Baceta J.I., Bernaola-Bilbao G., Nunez-Betelu K.,
Orue-Etxebarría X., Payros A., Pujalte V., Robin E. & Rocchia
R. 1997: Analysis of uppermost Cretaceous—lowermost Tertia-
ry hemipelagic successions in the Basque Country (western
Pyrenees): evidence for a sudden extinction of more than half
planktic foraminifer species at the K/T boundary. Bull. Soc.
Géol. France 168, 783—793.
Arenillas I., Arz J.A. & Molina E. 2004: A new high-resolution
planktic foraminiferal zonation and subzonation for the lower
Danian. Lethaia 37, 79—95.
Bubík M., Adamová M., Bąk M., Franců E., Franců J., Gedl P.,
Mikuláš R., Švábenická L. & Uchman A. 2002: Results of the
investigations at the Cretaceous/Tertiary boundary in the
Magura Flysch near Uzgruň. Geol. Výz. na Moravě a ve
Slezsku v roce 2001, Brno,18—22 (in Czech).
Chira C.M., Balc R., Cetean C., Juravle D.T., Filipescu S., Igritan A.,
Florea F. & Popa M.V. 2009: Cretaceous/Paleogene boundary in
north-eastern Romania. Bericht. Geol. Bundestanst. 78, 8.
Egger H., Rögl F. & Wagreich M. 2004: Biostratigraphy and facies
of Paleogene deep-water deposits at Gams (Gosau Group, Aus-
tria). Ann. Naturhist. Mus. Wien 106A, 281—307.
Egger H., Koeberl C., Wagreich M. & Stradner H. 2009: The Creta-
ceous-Paleogene (K/Pg) boundary at Gams, Austria: Nanno-
plankton stratigraphy and geochemistry of a bathyal
northwestern Tethyan setting. Stratigraphy 6, 333—347.
Gasiński M.A. 1997: Tethyan-Boreal connection: influence on the
evolution of mid-Cretaceous planktonic foraminiferids. Creta-
ceous Res. 18, 505—514.
Gasiński M.A. & Uchman A. 2009: Latest Maastrichtian foramin-
iferal assemblages from the Husów region (Skole Nappe, Out-
er Carpathians, Poland). Geol. Carpathica 60, 4, 283—294.
Gasiński M.A., Jugowiec M. & Ślączka A. 1999: Late Cretaceous
foraminiferids and calcareous nannoplankton from the Weglów-
ka Marls (Subsilesian Unit, Outer Carpathians, Poland). Geol.
Carpathica 50, 63—73.
Gasiński M.A., Leśniak T. & Piotrowski M. 2001: Latest Maas-
trichtian Foraminiferal Abathomphalus mayaroensis Zone in
the Subsilesian Unit (Polish Outer Carpathians). Bull. Polish
Acad. Sci. Earth Sci. 49, 89—97.
Gradstein F., Ogg J. & Smith A. 2004: A Geologic Time Scale.
Cambridge University Press, Cambridge, 1—589.
Kaiho K. & Lamolda M.A. 1999: Catastrophic extinction of plank-
tonic foraminifera at the Cretaceous-Tertiary boundary evi-
denced by stable isotopes and foraminiferal abundance at
Caravaca, Spain. Geology 27, 355—358.
Keller G. 2004: Paleoecology of Late Maastrichtian-early Danian
planktonic foraminifera in the eastern Tethys. J. Foram. Res.
34, 1, 49—73.
Kotlarczyk J. 1978: Stratigraphy of the Ropianka Formation or of
Inoceramian Beds in the Skole Unit of the Flysch Carpathians.
Prace Geol., Polska Akademia Nauk, Oddział w Krakowie, Ko-
misja Nauk Geologicznych 108, 1—82 (in Polish with English
summary).
Kotlarczyk J. 1988: Outline of the stratigraphy of the marginal tec-
tonic units of the Carpathian orogen. In: Kotlarczyk J., Pękala
K. & Gucik S. (Eds.): Przewodnik 59 Zjazdu Polskiego To-
warzystwa Geologicznego, Karpaty Przemyskie, 16—18
września 1988. Wydaw. AGH, Kraków, 23—62 (in Polish only).
Lahodynsky R. 1988: Lithostratigraphy and sedimentology across
the Cretaceous/Tertiary boundary in the Flyschgosau (Eastern
Alps, Austria). Riv. Esp. Paleont. N° Extr., 73—82.
MacLeod N. & Keller G. 1996: The Cretaceous-Tertiary Mass ex-
tinction: Biotic and environmental events. Norton Press, New
York, 1—595.
Melinte M.C. 1999: Cretaceous/Tertiary boundary in the East Car-
pathians (Romania), based on nannofloral evidence. Acta Pale-
ont. Romaniae 2 (1999), 269—273.
Molina E., Alegret L., Arenillas I., Arz J.A., Gallala N., Hardenbol
J., Salis K., von Steurbaut E., Vandenberghe N. & Zaghbib-
Turki D. 2006: The global boundary stratotype section and
point for the base of the Danian stage (Paleocene, Paleogene,
“Tertiary” Cenozoic) at El Kef, Tunisia – Original definition
and revision. Episodes 29, 263—273.
Olsson R.K., Hemleben C., Berggren W.A. & Huber B.T. 1999: At-
las of Paleocene planktonic foraminifera. Smithsonian Contr.
Paleobiol. 85, 1—252.
Peryt D., Lahodynsky R. & Durakiewicz T. 1997: Deep-water ag-
glutinated foraminiferal changes and stable isotope profiles
across the Cretaceous-Paleogene boundary in the Rotwandgra-
ben section, Eastern Alps (Austria). Palaeogeogr. Palaeocli-
matol. Palaeocol. 132, 287—307.
Premoli-Silva I. & Verga D. 2004: Practical manual of Cretaceous
Planktonic Foraminifera. International School on Planktonic
Foraminifera. 3° Course: Cretaceous. Verga et Rettori eds.
Universites of Perugia and Milano, Tipografia Pontfelcino,
Perugia, 1—283.
Rajchel J. 1990: Lithostratigraphy of the Upper Paleocene and
Eocene deposits in the Skole Unit. Zesz. Nauk. AGH, Geol. 48,
1—112 (in Polish with English summary).
Robaszynski F. & Caron M. 1995: Foraminiferes planctoniques du
Crétacé: commentaire de la zonation Europe-Méditerrané.
Bull. Soc. Géol. France 6, 681—692.
Wdowiarz S. 1949: Structure géologique des Karpates Marginales
au sud—est de Rzeszów. Biul. Państw. Inst. Geol. 11, 1—51.