GeolCarp_Vol62_No3_211

www.geologicacarpathica.sk

GEOLOGICA CARPATHICA

, JUNE 2011, 62, 3, 211—231 doi: 10.2478/v10096-011-0018-7

A late Burdigalian bathyal mollusc fauna from the Vienna

Basin (Slovakia)

MATHIAS HARZHAUSER

, OLEG MANDIC

and JAN SCHLÖGL

Natural History Museum Vienna, Burgring 7, A-1010 Vienna, Austria; mathias.harzhauser@nhm-wien.ac.at; oleg.mandic@nhm-wien.ac.at

Department of Geology and Paleontology, Faculty of Sciences, Comenius University, Mlynská dolina, pav. G, SK-842 15 Bratislava,

Slovak Republic; schlogl@fns.uniba.sk

(Manuscript received September 7, 2010; accepted in revised form November 9, 2010)

Abstract: This is the first record of a bathyal mollusc fauna from the late Early Miocene of the Central Paratethys. The
assemblage shows clear affinities to coeval faunas of the Turin Hills in the Mediterranean area and the Aquitaine Basin
in France. The overall biostratigraphic value of the assemblage is hard to estimate due to the general very poor knowl-
edge of Miocene bathyal faunas. Several species, however, are known from deep water deposits of the Middle Miocene
Badenian stage as well. This implies Early Miocene roots of parts of the Middle Miocene deep water fauna and suggests
a low turnover for bathyal mollusc communities at the Early-Middle Miocene boundary. The nassariid gastropod Nassarius
janschloegli Harzhauser nov. sp. and the naticid gastropod Polinices cerovaensis Harzhauser nov. sp. are introduced as
new species.

Key words: Early Miocene, Central Paratethys, Vienna Basin, bathyal, molluscs.

Introduction

The molluscs were collected at Cerová-Lieskové in the Slovak
Republic.  The  outcrop  is  situated  at  the  western  slope  of  the
Malé Karpaty Mts which forms the eastern margin of the cen-
tral Vienna Basin (Fig. 1A,B). The deposits of the former clay
pit are massive, locally laminated calcareous clay and clayey
silt  with  thin  tempestites  (up  to  5 mm  thick)  with  plant  re-
mains and several thin sandstone layers in the uppermost part
of the section (Fig. 1C) belonging to the Lakšárska Nová Ves
Formation (Špička & Zapletalová 1964). The grey sediments
have a beige-whitish weathering colour and are characterized
by a high amount of diatom skeletons. A detailed description
of the geological setting and the sedimentology will be given
elsewhere  (Hyžný  &  Schlögl  2011).  The  age  of  the  Cerová
section is Late Karpatian according to the regional stratigraphic
scheme, corresponding to the late Burdigalian.

Bathyal faunas in the Central Paratethys

The deep water fauna of the Paratethys is poorly known. In

the synthetic lists on Paratethyan Oligocene and Miocene gas-
tropods in Harzhauser & Piller (2007) most taxa derive from
shallow  marine  sections.  Shallow  bathyal  communities  from
the Late Oligocene of Eger in Hungary were described as Hinia-
Cadulus community by Báldi (1973). From Eggenburgian and
Ottnangian deposits, no bathyal mollusc faunas are known so
far.  The  so-called  Ottnangian  schlier  fauna  as  described  by
Hoernes  (1875)  was  sometimes  considered  to  have  lived  in
deeper marine settings but turned out to represent moderately
deep  sublittoral  environments  between  100—150 m  water
depth  (Grunert  pers.  comm.).  Information  on  the  Karpatian
mollusc fauna is based largely on littoral to shallow sublittoral

assemblages from mud-flats, sandy shores, lagoons and man-
groves  (Harzhauser  2002,  2003).  Nevertheless,  deeper  water
settings were widespread during Karpatian times. Spezzaferri
&  Coric  (2001)  and  Spezzaferri  et  al.  (2004)  discuss  water
depths of ca. 400—500 m in the Styrian Basin and of ca. 200 m
in the North Alpine Foreland Basin. One of their key-sections
is the middle Karpatian section Laa in Lower Austria which is
only  ca.  80 km  west  of  the  coeval  Cerová-Lieskové  section.
The mollusc faunas of the Austrian sections are very poor con-
sisting mainly of scattered shells of the cephalopod Aturia and
the pteropod Vaginella. The absence of a benthic mollusc fau-
na  may  be  related  to  the  dysoxic  conditions  on  the  sea  floor
(Spezzaferri et al. 2004).

During the Middle Miocene, deeper marine faunas are only

known  from  the  Badenian  basinal  clays.  The  water  depth  in
the Vienna Basin, where most of the classical Early to Middle
Badenian  faunas  derive  from,  ranged  around  300  meters
(Hohenegger et al. 2008). Similar conditions might have been
established  in  the  Transylvanian  Basin  where  deeper  marine
mollusc  assemblages  are  known  from  Lapugiu  de  Sus  and
Kostej.  Bathyal  associations,  however,  are  undescribed.  A
deep  sublittoral  mollusc  assemblage  from  the  Late  Badenian
of Devínska Nová Ves is the latest of its kind in the Central
Paratethys (Tomašových 1998).

Composition and biostratigraphy

The mollusc assemblage consists of 14 bivalves, 15 benthic

and 1 pelagic gastropods and 3 scaphopods. More than 85 %
of the benthic gastropods are carnivores, scavengers or para-
sites. In total numbers, the carnivorous tonnoideans, naticids
and conaceans predominate along with nassariids which are
scavengers and/or predators. Herbivores such as Calliotropis?

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sp. are extremely rare. Among the bivalves, two carnivorous
(Parvamussium and Cardiomya), four chemosymbiotic (Luci-
noidea  and  Solemya),  four  detritus  (Nuculoidea  and  Tellin-
idae)  and  four  suspension  feeding  (others)  bivalves  are
represented.  Such  composition  indicates  a  deposition  in  the
aphytal zone and a low contribution by transported taxa from
shallower  settings.  The  endemism  is  seemingly  high,  with  3
species  (18 %)  among  the  gastropods  and  4  species  (29 %)
among the bivalves, but should be considered with care in re-
spect to our poor knowledge on Miocene deep water faunas.
Within  the  scaphopods,  no  endemism  is  observed  as  all  taxa
are also known from the Mediterranean area.

The biostratigraphic value of the assemblage is hard to

evaluate due to the scarceness of ecologically equivalent fau-
nas.  Only  the  pteropod  Balantium  collina  (Janssen  &  Zorn
2001)  is  known  so  far  only  from  the  Burdigalian  of  Italy.
Calliotropis? sp., Polinices cerovaensis nov. sp. and Nassarius
janschloegli nov. sp. are only known from Cerová and may
represent  Karpatian  marker  species.  Amalda  glandiformis
(Lamarck,  1810),  Galeodea  echinophora  (Linnaeus,  1758),

Conolithus antidiluvianus (Brugui

re, 1792), Ringicula mi-

nor  (Grateloup,  1838)  and  Fissidentalium  badense  (Partsch
in  Hörnes,  1856)  appear  already  during  the  Early  Miocene
and persist throughout the Miocene. Others, such as Stellaria
testigera (Bronn, 1831), Mitrella hilberi (Cossmann, 1901),
Genota  valeriae  (Hoernes  &  Auinger,  1891),  Cylichna  cf.
salbriacensis  (Peyrot,  1932),  Sabatia  callifera  Boettger,
1906, Gadila gracilina Sacco, 1897 and Gadilina taurogra-
cilis Sacco, 1897 have not been known so far from the Early
Miocene but are documented from the Langhian. This indi-
cates that the Langhian deep water fauna is largely rooted in
Burdigalian  species.  Moreover,  the  herein  described
Burdigalian assemblage displays several – probably closely
related – counterparts in the Late Oligocene bathyal faunas
of  Hungary.  These  counterparts  are  Nassarius  schlotheimi
(Beyrich, 1854), Cylichna burdigalensis (sensu Báldi 1973),
Ringicula  paulucciae  (sensu  Báldi  1973),  Gadila  gracilina
(sensu  Báldi  1973),  Gadilina  taurogracilis  (sensu  Báldi
1973).  Thus,  no  big  turnover  seems  to  have  taken  place  as
far as this can be judged from the fragmentary data.

Fig. 1. A—B – Geographic position of the Cerová section on the eastern border of the Vienna Basin; 1 – European platform, 2 – Car-
pathian-Alpine externides, 3 – Pieniny Klippen Belt, 4 – Alpine-Carpathian-Dinaride and Pannonian internids, 5 – Neogene volcanites,
6 – Neogene basins. B – Bükk, CEA – Central Eastern Alps, CWC – Central Western Carpathians, M – Mecsek, MA – Magura
Group, NCA – Northern Calcareous Alps, RF – Rhenodanubian Flysh Zone, TCR – Transdanubian Central Range, W – Waschberg
Unit, ZD – Ždánice Unit. C – lithology of the section; numbers indicate the position of the samples as mentioned in the text; a – mas-
sive calcareous clay, b – thin tempestite layers with plant remains, c – thin siltstone/sandstone layers or silt lenses.

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This pattern is also reflected by the bivalve assemblage.

All bivalve species from Cerová pass the Early-Middle Mio-
cene boundary and are present also in the Badenian deep wa-
ter  deposits.  Five  species  are  restricted  to  the  Early  and
Middle  Miocene:  Leionucula  ehrlichi  (Hoernes,  1875),
Limaria  labani  (Meznerics,  1936),  Lucina  callipteryx
Tournouer,  1874,  Laternula  fuchsi  (Hoernes,  1875),  and
Cardiomya elegantissima (Hoernes, 1875). All other species
except for Nucula mayeri (Hörnes, 1865) persist even into the
Pliocene.  The  latter,  together  with  Yoldia  nitida  (Brocchi,
1814),  Solemya  doderleini  Mayer,  1861,  Atrina  pectinata
(Linnaeus,  1767),  Macoma  elliptica  (Brocchi,  1814),  and
Thyasira flexuosa (Montagu, 1803), already occurs during the
Oligocene.

Bathymetry

An ecologically highly significant taxon is the scaphopod

Gadilina  taurogracilis.  Its  descendent  Gadilina  triquetra
(Brocchi,  1814)  is  widespread  in  Pliocene  deposits  of  Italy.
According  to  Ceregato  et  al.  (2007)  it  is  strictly  bathyal  and
indicative  for  unstable  deep  marine  environments  with  high
sedimentation  rates.  Such  Gadilina-dominated  assemblages
are part of the widespread Early Pliocene Korobkovia oblon-
ga—Jupiteria  concava  paleocommunity  of  Ceregato  et  al.
(2007)  which  contains  also  Stellaria  testigera  and  Galeodea
echinophora.  As  mentioned  above,  the  Hungarian  Hinia-
Cadulus  paleocommunity  of  Báldi  (1973)  is  an  Oligocene
counterpart  in  the  Paratethys  (=Nassarius-Gadila  paleocom-
munity according to modern systematics).

Material

The material is stored in the collection of the Natural Histo-

ry Museum Bratislava, Department of Geology and Paleonto-
logy, Faculty of Natural Sciences, Comenius University in

Bratislava and in the Natural History Museum Vienna (only
type material).

Class: Gastropoda Cuvier, 1797

Subclass: Orthogastropoda Ponder & Lindberg, 1996

Superorder: Vetigastropoda Salvini-Plawen & Haszprunar,

1987

Superfamily: Seguenzioidea Verrill, 1884

Family: Chilodontidae Wenz, 1938

?Genus: Calliotropis Seguenza, 1903

Calliotropis

? sp.

M a t e r i a l: 1 shell fragment (sample number: 20—21); dia-

meter: 5.3 mm.

R e m a r k s: A very poorly preserved specimen, which does

not allow a clear identification. The base bears 4—5 blunt spi-
ral  ribs  which  are  crossed  by  sharp,  wide-spaced  axial  ribs.
Small  nodes  appear  at  the  intersections  and  a  slightly  more
prominent spiral rib forms a peripheral keel. The umbilicus is
wide  and  open.  No  comparable  vetigastropod  is  described
from Paratethyan and Mediterranean Miocene deposits.

Suborder: Hypsogastropoda Ponder & Lindberg, 1997

Infraordo: Littorinimorpha Golikov & Starobogatov, 1975

Superfamily: Naticoidea Guilding, 1834

Family: Naticidae Guilding, 1834

Subfamily: Polinicinae Gray, 1847

Genus: Polinices Montfort, 1810

Polinices cerovaensis

Harzhauser nov. sp.

Fig. 2.1—3

M a t e r i a l: 12 shells (sample numbers: 5—6, 6, 7, 7—8, 13—

14, 14, 15, 15—16, 16—17, 19, 20, 21).

H o l o t y p e: NHMW 2010/0080/0001; height: 6.9 mm,

diameter: 6.5 mm (Fig. 2.1).

Fig. 2. 1—3 – Polinices cerovaensis Harzhauser nov. sp. 1 – holotype NHMW 2010/0080/0001, 2 – paratype NHMW 2010/0080/0002,
3 – a third, not fully grown specimen.

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P a r a t y p e: NHMW 2010/0080/0002; height: 9.1 mm, dia-

meter: 7 mm (Fig. 2.2).

S t r a t u m t y p i c u m: Grey calcareous silt of the Lakšárska

Nová Ves Formation.

T y p e l o c a l i t y: Cerová-Lieskové, Slovak Republic.
A g e: Early Miocene, late Burdigalian; Karpatian.
N a m e: Referring to the type locality.
D i a g n o s i s: A small Polinices with globose to broad drop-

shaped outline. The small, moderately thickened parietal cal-
lus bears a shallow sulcus.

D e s c r i p t i o n: A small naticid which ranges between 5—

10 mm in height. Small, depressed turbiniform protoconch of
1.7 convex whorls. Globose to slender globose shell consist-
ing  of  3—4  convex  whorls.  The  spire  height  is  quite  variable
and ranges from moderately elevated to low. The last whorl is
nearly  straight  sided  close  to  the  upper  suture  or  may  even
form a very shallow and indistinct concavity, causing a broad
drop-shaped  outline.  Shell  surface  smooth  except  for  proso-
cline growth lines. These may be more prominent in the adapi-
cal third of the whorls. In one specimen these growth lines are
quite strongly developed on the penultimate whorl. The parie-
tal callus is small, moderately thick, well demarcated from the
base and develops a shallow sulcus in its middle. This callus
covers large parts of the umbilicus which is deep but narrow
and sickle shaped.

R e m a r k s: A common species at Cerová. Its small size

and the sulcus on the parietal callus separate this species dis-
tinctly  from  Polinices  pseudoredemptus  (Friedberg,  1923)
which  occurs  in  coastal  marine  and  brackish  environments
during  the  Karpatian  and  Badenian  (Harzhauser  2002).  Po-
linices  proredemptus  (Sacco,  1891)  from  the  Early  Miocene
of Italy and France, lacks the sulcus, has a depressed spherical
shape  and  develops  a  wide  lobe  of  the  parietal  callus  and  a
second lobe on the base (Cossmann & Peyrot 1919). The sul-
cus also separates the new species from the French Early Mio-
cene Polinices benoisti (Cossmann & Peyrot, 1919) which has
a  similar  shape  but  develops  a  wider  and  deeper  umbilicus
(Cosmann  &  Peyrot  1919;  Lozouet  et  al.  2001).  Polinices
turbinoides  (Grateloup,  1828),  from  the  Early  Miocene  of

France, develops a more slender shape with high penultimate
whorl (Cossmann & Peyrot 1919).

Family: Xenophoridae Troschel, 1852

Genus: Stellaria Schmidt in Möller, 1832

Stellaria testigera

(Bronn, 1831)

Fig. 3.2—4

1831 Phos testigerus Bronn, p. 61
2004 Stellaria testigera testigera (Bronn, 1831) – Landau et al., p. 85,

pl. 19, figs. 3—4 (cum syn.)

M a t e r i a l: 2 shells (sample numbers: 15, 20); diameter:

27—30 mm.

R e m a r k s: A rare species at Cerová. The identification is

based  on  the  characteristic  rugose  sculpture  that  undulates
along the lower sutures and especially close to the long digita-
tions. The attached objects are small molluscs such as nassari-
ids  and  bivalve  fragments.  This  is  the  first  Early  Miocene
record  of  Stellaria  testigera  which  is  otherwise  documented
only  from  the  Middle  Miocene  to  Pliocene  of  the  Paratethys
and the Mediterranean Sea (see Landau et al. 2004 for details).
In the Vienna Basin it is restricted to deeper marine clays of
the Badenian stage.

Superfamily: Tonnoidea Suter, 1913

Family: Tonnidae Suter 1913

Family: Cassidae Latreille, 1825

Genus: Galeodea Link, 1807

Galeodea echinophora

(Linnaeus, 1758)

Fig. 3.1

1758 Buccinum echinophorum Linnaeus, p. 735
2009 Galeodea echinophora (Linnaeus, 1758) – Landau, p. 66, pl. 3,

figs. 1—2 (cum syn.)

M a t e r i a l: 17 shells (sample numbers: 5—6, 15, 16—17,

17—18, 18—19, 20—21, P2—5); mostly fragments; the size
seems to have ranged between 40—50 mm.

Fig. 3. 1 – Galeodea echinophora (Linnaeus, 1758), 2—4 – Stellaria testigera (Bronn, 1831).

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R e m a r k s: One of the most abundant gastropods at

Cerová. A strongly sculptured morphotype predominates with
three spiral rows of strong nodes on the last whorl and a fourth
row of spiny nodes along the shoulder. The shells are usually
strongly  fragmented  and  have  been  destroyed  before  deposi-
tion. Obviously, it was the favourite prey for crushing preda-
tors  such  as  decapods  or  molluscivore  fish.  The  species
appears during the Early Miocene and is still present in the
Mediterranean  Sea  and  along  the  coast  of  western  Africa
(Ardovini & Cossignani 2004). In the Paratethys it is known
from  the  Eggenburgian  to  the  Badenian  (see  Landau  et  al.
2009 for details).

Infraorder: Neogastropoda Wenz, 1938

Superfamily: Buccinoidea Rafinesque, 1815

Family: Columbellidae Swainson, 1840

Genus: Mitrella Risso, 1826

Mitrella hilberi

(Cossmann, 1901) nov. comb.

Fig. 4.2

1879 Columbella carinata Hilber, p. 6, pl. 1, fig. 3 (non Columbella cari-

nata Hinds, 1844)

1880 Columbella carinata Hilber – Hoernes & Auinger, p. 97, pl.12,

figs. 9—11

1901 Atilia (Macrurella) hilberi Cossmann, p. 245
1966 Columbella (Atilia) hilberi Cossmann – Strausz, p. 293, pl. 12,

fig. 12

M a t e r i a l: 1 cast and silicone mould (sample number: 17);

height: 15.4 mm, diameter: 4.3 mm.

R e m a r k s: The specimen differs from the type of Mitrella

hilberi from the Early Badenian of the Styrian Basin in its less
angulated transition towards the base. Specimens from the
Early Badenian of Kostej in Romania, illustrated by Hoernes
& Auinger (1880), are also characterized by a less prominent
angulation and correspond fully to the shell from Cerová. The
specimen from the Badenian of Sámsonháza in Hungary, il-

lustrated  in  Strausz  (1966),  also  belongs  to  this  morphotype.
The  separation  from  Mitrella  petersi  (Hilber,  1879)  would
need  further  confirmation  but  seems  to  be  justified  based  on
the nearly straight sided whorls and the more slender outline
of  the  type  of  M.  petersi.  Mitrella  aquitanica  (Peyrot,  1925)
from the Aquitanian of France is stout and has a shorter last
whorl.

Mitrella hilberi was known so far only from marl and clay

of the Badenian stage.

Family: Nassariidae Iredale, 1916

Genus: Nassarius Duméril, 1806

Nassarius janschloegli

Harzhauser nov. sp.

Fig. 5.1—4

M a t e r i a l: 22 shells (sample numbers: 5—6, 7, 7—8, 8—9, 12,

13—14, 14, 15, 16—17, 17, 17—18, 18—19, 19—20, 21, 21—22,
P2—5).

H o l o t y p e: NHMW 2010/0079/0001; height: 6.1 mm, dia-

meter: 4.1 mm (Fig. 5.1).

P a r a t y p e 1: NHMW 2010/0079/0002; height: 6.9 mm,

diameter: 4.3 mm (Fig. 5.2).

P a r a t y p e 2: NHMW 2010/0079/0003; diameter: 3.3 mm

(Fig. 5.4).

S t r a t u m t y p i c u m: Grey calcareous silt of the Lakšárska

Nová Ves Formation.

T y p e l o c a l i t y: Cerová-Lieskové, Slovak Republic.
A g e: Early Miocene, late Burdigalian, Karpatian.
N a m e: In honor of Jan Schlögl, paleontologist at the

Comenius University, Bratislava.

D i a g n o s i s: Small shell with elevated spire of 2.5 weakly

convex spire whorls and a broader, convex last whorl. Sculp-
ture consisting of prosocline to slightly sigmoidal axial ribs
which cross weaker, broad spiral ribs. Nodes appear at the in-
tersections. A conspicuous row of nodes at the upper suture
causes a stepped outline.

Fig. 4. 1 – Amalda glandiformis (Lamarck, 1810), 2 – Mitrella hilberi (Cossmann, 1901), 3—4 – Genota valeriae (Hoernes & Auinger,
1891), 5 – Conolithus antidiluvianus (Brugui

re, 1792).

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D e s c r i p t i o n: The species has a 1-mm-broad, low dome-

shaped protoconch consisting of 3.5 convex whorls. The sec-
ond  protoconch  whorl  bears  2—3  faint  spiral  threads  in  the
upper  third;  additional  wrinkled  spirals  seem  to  appear  close
to the lower suture on the last protoconch whorl (this feature is
obscured by the poor preservation). The teleoconch develops a
moderately  high  and  stepped  spire  of  three  weakly  convex
whorls and a broader, convex last whorl. The sculpture of the
early whorls consists of narrow prosocline to weakly sigmoi-
dal axial ribs and less prominent, broader spiral ribs separated
by  shallow  interspaces.  The  intersections  tend  to  form  small
pointed  nodes.  The  spiral  ribs  become  stronger  towards  the
last whorl but the axial ribs remain the dominating sculpture.
The uppermost row of nodes along the upper suture is distinct-
ly  stronger,  slightly  separated  from  the  next  row  and  causes
the stepped profile. In some specimens, the second row below
the upper suture is also somewhat more prominent and forms
a  pair  of  nodes  on  the  adapical  termination  of  the  axial  ribs.
The spiral sculpture becomes weak on the last whorl below the
point of maximum convexity but reappears as 2—3 sharp bead-
ed spiral threads on the base. The aperture is ovoid; columella
strongly  concave,  adapically  nearly  straight  with  two  broad
denticles.  Two  further  denticles  occur  close  to  the  wide  and
deep siphonal canal. Columellar callus moderately thickened,
clearly delimited. Outer lip terminating in a smooth bevelled
edge;  five  strong  and  elongate  denticles  occur  at  some  dis-
tance from the termination.

R e m a r k s: A highly reminiscent species is Nassarius

schlotheimi  (Beyrich,  1854)  as  described  from  the  Late
Oligocene  of  Hungary  (Báldi  1973).  It  displays  a  similar
variability in sculpture, agrees in the development of the adsu-
tural row of nodes and has a similar large protoconch but lacks
the prominent denticles in the outer lip and has more convex
whorls. Interestingly, Nassarius schlotheimi was described by
Báldi  (1973)  as  member  of  the  bathyal  communities.  Both
species belong to a morphological group which is represent-
ed  during  the  Early  Miocene  by  species  such  as  Nassarius
pauli  (Hoernes,  1875),  Nassarius  illovensis  (Hoernes  &
Auinger  1882),  Nassarius  perpulchra  (Bellardi,  1882)  and
Nassarius incerta (Bellardi, 1882).

Nassarius pauli, from the Ottnangian stage of Upper Austria,

might be a close relative. Parallels are the large bulbous proto-
conch  and  the  row  of  small  nodes  along  the  shoulder
(Harzhauser & Kowalke 2004). Differences are the much larger
size of N. pauli, which attains about double the height, and the
much shorter spire. Moreover, N. pauli develops broader axial
ribs  which  bear  broader  rounded  nodes.  Nassarius  illovensis,
which occurs in Middle Miocene offshore marls in the Para-
tethys,  is  much  larger  and  stout  (Harzhauser  &  Kowalke
2004). Its spire whorls are more convex; the sculpture differs
in  its  dense  and  regular  pattern  of  nodes.  Nassarius  incerta,
from  the  Burdigalian  of  the  Italian  Turin  Hills,  is  strongly
reminiscent of the species from Cerová but lacks the stepped
outline, the adsutural row of nodes and is much larger (Ferrero
Mortara et al. 1981). Nassarius perpulchra, from the Burdiga-
lian of the Italian Turin Hills, develops a very similar sculp-
ture but has a broader spire and develops a conspicuous bulgy
convexity in the lower third of the last whorl (Ferrero Mortara
et  al.  1981).  Nassarius  janschloegli  was  a  small  nassariid
which was probably adapted to deeper marine environments.

Superfamily: Olivoidea Latreille, 1825

Family: Olividae Latreille, 1825

Genus: Amalda Adams & Adams, 1852

Amalda glandiformis

(Lamarck, 1810)

Fig. 4.1

1810 Ancillaria glandiformis Lamarck, p. 305
1997 Ancilla (Baryspira) glandiformis (Lamarck) – Bałuk, p. 24, pl. 6,

figs. 1—11

2002 Amalda (Baryspira) glandiformis (Lamarck) – Harzhauser, p. 109,

pl. 8, fig. 19 (cum syn.)

M a t e r i a l: 1 spire fragment (found in scree material); di-

ameter: 9 mm.

R e m a r k s: The wide apical angle and thick callus agree

fully with Amalda glandiformis and can be easily separated
from Amalda obsoleta (Brocchi, 1814), which is typical for
deeper marine deposits. Typically, the slender elongate A. ob-
soleta predominates in the Badenian clay of the Vienna Basin

Fig. 5. 1—4 – Nassarius janschloegli Harzhauser nov. sp., 1 – holotype NHMW 2010/0079/0001, 2 – paratype NHMW 2010/00879/0002,
4 – paratype NHMW 2010/00879/0003.

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and  the  Korytnica  clay  in  Poland,  whilst  small-sized  A.
glandiformis  are  rare.  This  polymorphic  species  (or  species
flock)  appears  during  the  Oligocene,  flourishes  during  the
Miocene  in  all  European  seas  and  persists  in  the  Mediterra-
nean Sea and the Eastern Atlantic up the Pliocene (Landau &
Marques da Silva 2006). During the Karpatian it is a frequent
species in shallow marine settings of the Korneuburg Basin in
Austria (Harzhauser 2002).

Superfamily: Conoidea Fleming, 1822

Family: Conolithidae Tucker and Tenorio, 2009

Genus: Conolithus Swainson, 1840

Conolithus antidiluvianus

(Brugui

re, 1792)

Fig. 4.5

1792 Conus antidiluvianus Brugui

re, p. 637, pl. 347, fig. 6

1964 Conus antidiluvianus Brugui

re – Hall, p. 17, pl. 2, fig. 7

1966 Conus (Conolithus) antediluvianus Brugui

re – Strausz, p. 451,

pl. 66, fig. 10, pl. 67, fig. 1

1998 Conus (Conolithus) antediluvianus Brugui

re – Schultz, p. 72,

pl. 29, fig. 9

M a t e r i a l: 1 shell (found in scree material); height: 8 mm,

diameter: 3.5 mm.

R e m a r k s: The small slender shell is characterized by a

stepped  spire  with  small,  triangular  nodes  at  the  shoulder,
pointing in adapical direction. The base bears a sculpture of
delicate spiral furrows in the lower third. These features are
typical  for  Conolithus  antidiluvianus,  which  is  a  common
species in Miocene offshore clays. In the Paratethys, it is re-
corded from Ottnangian “Schlier” facies in the North Alpine
Foreland  Basin  (Hoernes  1875),  from  the  Karpatian  of  the
Vienna  Basin  (Harzhauser  2003)  and  from  the  Badenian  of
the Vienna Basin and the Pannonian Basin complex (Strausz
1966).  In  the  Mediterranean  and  the  Eastern  Atlantic  it  ap-
pears  during  the  Early  Miocene  (e.g.  late  Burdigalian  deep
sublittoral environments of the Mut Basin in Turkey; Mandic
et  al.  2004),  reaches  the  North  Sea  during  the  Middle  Mio-

cene (Janssen 1984) and persists in the Mediterranean Sea up
to the Pliocene (Chirli 1997).

Hall (1964) pointed out that antediluvianus, which is com-

monly used in the literature, is a spelling error for the correct
antidiluvianus.

Family: Conorbinae de Gregorio, 1890

Genus: Genota Adams & Adams, 1853

Genota valeriae

(Hoernes & Auinger, 1891)

Fig. 4.3—4

1891 Pleurotoma (Genota) valeriae Hoernes & Auinger, p. 311, pl. 34,

fig. 15

1966 Genota ramosa valeriae Hoernes & Auinger – Strausz, p. 448,

fig. 198

2003 Genota (Genota) valeriae (Hoernes & Auinger, 1891) – Bałuk,

p. 55, pl. 18, figs. 4—6

M a t e r i a l: 2 shells (sample numbers: 15, 17); height: ca.

30 mm, diameter: ca. 9 mm.

R e m a r k s: The fragments agree well with the type materi-

al from Lapugiu de Sus in Romania in the collections of the
Natural History Museum Vienna. Its protoconch corresponds
largely to that of Genota ramosa (Basterot, 1825) as illustrat-
ed by Janssen (1984) supporting a close relationship between
the two species. Whilst G. ramosa is widespread in shallow
marine environments during the Early Miocene in the Mediter-
ranean, the Eastern Atlantic and the North Sea, its relative
Genota valeriae is known so far only from deeper marine Bade-
nian marls of Romania, Hungary, Poland and Austria. The oc-
currence at Cerová is thus the oldest record of the species.

Superfamily: Pyramidelloidea Gray, 1840

Family: Pyramidellidae Gray, 1840

Genus: Pyramidella Lamarck, 1799

Pyramidella

cf. gratteloupi d’Orbigny, 1852

Fig. 6.1

Fig. 6. 1 – Pyramidella cf. gratteloupi d’Orbigny, 1852, 2 – Ringicula minor (Grateloup, 1838), 3—4 – Balantium collina (Janssen &
Zorn, 2001).

è
è

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cf. 1852 Pyramidella gratteloupi d’Orbigny, p. 34
cf. 2001 Pyramidella gratteloupi (d’Orbigny) – Lozouet et al., p. 74,

pl. 36, fig. 6

1932 Roxania helvetica var. salbriacensis Peyrot, p. 198, pl. 13,

figs. 36, 40

M a t e r i a l: 1 shell (sample number: 16); height: 6.8 mm,

diameter: 2.1 mm.

R e m a r k s: The preservation does not allow a clear identi-

fication. Size and shape are reminiscent of the Early Miocene
Pyramidella gratteloupi from France. Shells from the Middle
Miocene of the Paratethys which are treated as Pyramidella
plicosa (Bronn, 1838) in the literature and in the collection of
the NHMW differ in their lower early spire whorls. The later
teleoconch of these shells, however, is nearly indistinguish-
able from the shell from Cerová. (Note that the original spell-
ing in d’Orbigny (1852) is gratteloupi instead of grateloupi.)

Order: Opistobranchia Milne-Edwards, 1848

Suborder: Cephalaspidea Fischer, 1883

Superfamily: Ringiculoidea Philippi, 1853

Family: Ringiculidae Philippi, 1853

Genus: Ringicula Deshayes in Lamarck, 1838

Ringicula minor

(Grateloup, 1838)

Fig. 6.2

1838 Auricula ringens var. b. minor Grateloup, p. 286, pl. 6, fig. 8
1878 Ringicula paulucciae Morlet, p. 266, pl. 6, fig. 6, pl. 8, fig. 9
1878 Ringicula (Ringiculella) tournoueri Morlet, p. 287, pl. 6, fig. 10
1954 Ringicula (Ringiculella) auriculata paulucciae Morlet – Berger,

p. 7, figs. 3—18

1984 Ringicula (Ringiculella) paulucciae Morlet – Švagrovský, p. 184,

pl. 2, fig. 6

1998 Ringicula paulucciae Morlet – Valdés & Héros, p. 700, fig. 2f
1998 Ringicula tournoueri Morlet – Valdés & Héros, p. 700, fig. 3g
2001 Ringicula minor (Grateloup) – Lozouet et al., p. 80, pl. 37, fig. 1

M a t e r i a l: 3 fragments (sample numbers: 13, 16, 19—20);

diameter: 5.1 mm.

R e m a r k s: The ringiculids of the Paratethys have been

studied  by  Berger  (1954)  who  mainly  referred  to  Morlet
(1878) but failed to integrate other French literature (e.g. Pey-
rot  1932).  Consequently,  he  assigned  the  Early  and  Middle
Miocene Paratethyan ringiculid shells with elongate spire and
spiral sculpture to Ringicula paulucciae Morlet, 1878 and its
synonym  Ringicula  tournoueri  Morlet,  1878  (illustrated  in
Valdez  &  Herós  1998).  Later,  Lozouet  et  al.  (2001)  showed
that the types of these species are conspecific with Ringicula
minor (Grateloup, 1838). Ringicula minor, though referred to
as  Ringicula  paulucciae  in  Paratethys  literature,  is  a  wide-
spread  species  during  the  Early  and  Middle  Miocene  in  the
Eastern Atlantic, the Mediterranean Sea and the Central Para-
tethys.  Many  Paratethyan  shells,  referred  to  as  Ringicula
buccinea (Brocchi, 1814), by Strausz (1966) and Atanacković
(1985) represent R. minor as well.

Superfamily: Philinoidea Gray, 1850

Family: Cylichnidae Adams & Adams, 1854

Genus: Cylichna Lovén, 1846

Cylichna

cf. salbriacensis (Peyrot, 1932)

Fig. 7.2—4

M a t e r i a l: 8 shells (sample numbers: 5—6, 7—8, 9, 16—17,

17—18, 19—20, 20—21, P2—5); height: 9—16 mm, diameter:
4.5—7 mm.

R e m a r k s: A large, subcylindrical shell with conspicuous

spiral sculpture on the spire and the lower third of the last
whorl. The prominent spirals on the base are duplex-rib pairs
with a narrow thread-like furrow separating each pair. These
are then separated from each other by deep spiral furrows. The
sculpture is only reduced in the middle part of the whorl.

Cylichna salbriacensis (Peyrot, 1932), from the Langhian

of  the  Aquitaine  is  highly  reminiscent  concerning  shape,
size  and  sculpture.  This  species  was  introduced  by  Peyrot
(1932)  as  a  subspecies  of  Roxania  helvetica  Peyrot,  1932
(non  Roxania  helvetica  Berger,  1953)  from  Saubrigues  in
France. Cylichna helvetica differs from  Cylichna salbriacen-
sis in its much finer sculpture and the elongate barrel-shaped
outline, whereas C. salbriacensis develops a slight convexity
on the adapical third similar to the shells from Cerová.

The species is missing in the revision on Miocene opistho-

branchs from the Vienna Basin by Berger (1953). This may be
caused by the fact that Berger (1953) did not consider any spe-
cies from the Miocene of France described by Peyrot (1932).
Nevertheless, a shell in the collection of the NHMW from the
Early Badenian of Kostej in Romania, which was mentioned
by  Hörnes  (1856)  as  Bulla  brocchi  Michelotti,  1847,  is  con-
specific  with  the  specimens  from  Cerová.  The  Romanian
shells were later treated as the much younger Retusa brocchii
(Michelotti) by Berger (1953) or as Retusa testiculina (Bonelli)
by Boettger (1906). The former name refers to a Late Miocene
or  Pliocene  Mediterranean  species  with  very  weak  to  absent
spiral sculpture and the latter name is a mere catalogue name
and is invalid. Shells from the Early Badenian of Lower Aus-
tria, erroneously identified as C. pliocrassa by Berger (1953),
may  be  conspecific  with  the  species  from  Cerová.  The
Pliocene Cylichnina pliocrassa Sacco, 1897, however, differs
from the Cerová species and that of Berger (1953) in its deli-
cate sculpture and the sub-parallel flanks of the last whorl.

Genus: Sabatia Bellardi, 1877

Sabatia callifera

Boettger, 1906

Fig. 7.1

1856 Bulla utriculus Brocchi – Hörnes, p. 618 (partim), pl. 50, fig. 2

(non Roxania utriculus (Brocchi, 1814))

1906 Sabatia callifera n. sp. Boettger, p. 205
1934 Sabatia callifera Boettger – Zilch, p. 278, pl. 22, fig. 20
1953 Roxania (Sabatia) callifera callifera (Boettger) – Berger, p. 112,

pl. 18, fig. 78

1985 Roxania (Roxania) callifera (Boettger) – Atanacković, p. 188,

pl. 42, figs. 4—5

M a t e r i a l: 4 specimens (sample numbers: 13, 14, 16—17,

19); height: 5 mm, diameter: 3 mm.

R e m a r k s: Only two fragments were found at Cerová.

Most parts of the shells are covered by sediment and therefore
the identification is difficult. Nevertheless, size, shape, spire

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morphology  and  the  conspicuous  sculpture  agree  fully  with
Middle Miocene shells of Sabatia callifera. This rare species
was described so far only from deeper marine clays of  Early
Badenian  age  from  Lapugiu  de  Sus  in  Romania  and  from  a
few sections in the Vienna Basin (Berger 1953). Additionally,
it  is  reported  from  the  Badenian  of  Bulgaria  and  Bosnia
(Atanacković  1985).  This  record  is,  thus,  the  oldest  occur-
rence of the species.

Superorder: Heterobranchia Gray, 1840

Order: Thecosomata Blainville, 1823

Superfamily: Cavoliniodea Fischer, 1883

Family: Cliidae Jeffreys, 1869

Genus: Balantium Bellardi, 1872

Balantium collina

(Janssen & Zorn, 2001)

Fig. 6.3—4

2001 Clio (Balantium) collina spec. nov. Janssen & Zorn, p. 47, fig. 1

M a t e r i a l: 12 shells, (sample numbers: 5—6, 6—7, 8—9,

10—11, 12, 13—14, 14—15, 16—17, 17—18, 19, 19—20, 21);
length: 16 mm, diameter: 8 mm.

R e m a r k s: A common species at Cerová; the shells appear

as  isolated  specimens  without  forming  distinct  layers.  The
species  was  originally  described  by  Janssen  &  Zorn  (2001)
from  the  Burdigalian  Termô-Fôr

Formation of the Turin

Hills. It is thus a good biostratigraphic marker for the Burdiga-
lian age of the Lakšárska Nová Ves Formation and documents
the open marine connection between the Karpatian Paratethys
and the Burdigalian Mediterranean Sea.

Class: Scaphopoda Bronn, 1862

Order: Dentaliida da Cosat, 1776

Family: Dentaliidae Children, 1834

Genus: Fissidentalium Fischer, 1885

Fissidentalium badense

(Partsch in Hörnes, 1856)

Fig. 8.6—7

1856 Dentalium Badense Hörnes, p. 652, pl. 50, fig. 30
1991 Fissidentalium badense (Partsch in Hörnes, 1856) – Pavia, p. 146,

pl. 5, fig. 4, pl. 6, fig. 6 (cum syn.)

M a t e r i a l: 5 shells, (sample numbers: 5—6, 6—7, 17, 18,

21); length: 41 mm, diameter: 5 mm.

R e m a r k s: This species appears during the Burdigalian in

the Mediterranean where it is found in the Turin Hills (Sacco
1897) and in deep water deposits of the Mut Basin in Turkey
(Mandic et al. 2004). The specimens from Cerová are the first
Early Miocene record of this species from the Paratethys. Dur-
ing the Langhian it is widespread from the Eastern Atlantic
(Cossmann & Peyrot 1917), the Mediterranean and the Para-
tethys Sea and persists in the Mediterranean Sea up the Late
Miocene (Pavia 1991).

Family: Gadilinidae Chistikov, 1975

Genus: Gadilina Foresti, 1895

Gadilina taurogracilis

Sacco, 1897

Fig. 8.1—2

1897 Gadilina triquetra var. taurogracilis Sacco, p. 114, pl. 10,

figs. 44—46

1991 Gadilina triquetra taurogracilis Sacco – Pavia, p. 130, pl. 7,

fig. 3a—b

M a t e r i a l : 8 shells, (sample number: 8—9, 10—11, 14,

16—17, 18—19, 19—20, P2—5 and scree material); length:
29 mm, diameter: 3 mm.

R e m a r k s: The slender, smooth and glossy shells are weak-

ly triangular in cross-section with rounded edges. This species
was introduced by Sacco (1897) from the Langhian of the Mon-
te dei Cappuccini in the Turin Hills. He considered it a subspe-
cies  of  the  Late  Miocene  to  Pliocene  deep  water  scaphopod
Gadilina  triquetra  (Brocchi,  1814).  Pavia  (1991)  designated  a
lectotype,  re-illustrated  the  specimen  and  emphasized  the  less
angular edges and the more slender outline of the Middle Mio-
cene taxon. An even earlier record of this species is presented

Fig. 7. 1 – Sabatia callifera Boettger, 1906, 2—4 – Cylichna cf. salbriacensis (Peyrot, 1932).

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by Cosmann & Peyrot (1917) from the Chattian of Aquitaine
in France and by Báldi (1973) from the Chattian of Hungary.
The small, less curved and ovoid instead of trigonal Oligocene
specimens, however, seem to represent another species.

Order: Gadilida Starobogatov, 1974

Suborder: Gadilimorpha Steiner, 1992

Family: Gadilidae Stoliczka, 1868

Genus: Gadila Gray, 1847

Gadila gracilina

Sacco, 1897

Fig. 8.3—5

1897 Gadila gadus var. gracilina Sacco, p. 117, pl. 10, figs. 86—87
1991 Gadila gracilina Sacco – Pavia, p. 144, pl. 8, figs. 1—4

M a t e r i a l: 8 shells, (sample numbers: 5—6, 6—7, 13—14,

15—16, 16—17, 17—18, 19—20, P2—5); length: 13—19 mm, dia-
meter: 3.5 mm.

R e m a r k s: The most common scaphopod at Cerová. The

maximum  diameter  of  the  smooth  and  glossy  shell  is  devel-
oped  close  to  the  rapidly  contracting  anterior  aperture.  This
feature distinguishes Gadila gracilina from the shorter Gadila
ventricosa  (Bronn,  1827)  which  has  a  much  longer  part  be-
tween aperture and maximum diameter. The latter species ap-
pears in the Badenian of the Paratethys (Pavia 1991) whereas
Gadila  gracilina  was  unknown  so  far  from  this  sea.  It  was
originally described by Sacco (1897) from the Langhian of the
Monte Cappuccini in the Turin Hills and from the Late Mio-
cene of Sant’Agata. One syntype was re-illustrated by Ferrero-
Mortara  et  al.  (1984)  and  Pavia  (1991)  provided  a
re-description. Another Langhian occurrence is mentioned by
Cosmann  &  Peyrot  (1917)  from  the  Aquitaine.  Chattian  oc-
currences  in  Hungary  (Báldi  1973)  and  France  (Cosmann  &
Peyrot 1917) need confirmation.

Class: Bivalvia Linnaeus, 1758

Subclass: Protobranchia Pelseneer, 1889

Order: Nuculida Dall, 1889

Superfamily: Nuculoidea Gray, 1824

Family: Nuculidae Gray, 1824

Genus: Nucula Lamarck, 1799

Nucula mayeri

Hörnes, 1865

Fig. 9.1

1865 Nucula Mayeri Hörn. – Hörnes, p. 296—297, pl. 38, fig. 1
1875 Nucula Mayeri M. Hoern. – Hoernes, p. 377, pl. 14, fig. 10
1912 Nucula Mayeri Hoernes – Cossmann & Peyrot, p. 219—221, pl. 5,

figs. 21—24

2001 Nucula (Nucula) mayeri Hörnes, 1865 – Schultz, p. 2, pl. 1,

figs. 3—4

M a t e r i a l: 1 single left shell fragment embedded in sedi-

ment with interior side down (sample 18). Length (interpo-
lated) 7.2 mm, height (interpolated) 6.4 mm.

R e m a r k s: Shell small, strongly convex, ovoid in outline,

elongated  anteriorly,  smooth  with  very  fine  concentric
lamellae  on  the  proximal  shell  exterior  and  very  fine  radial
striae of which proximally every fifth is a little more promi-
nent.  Determination  is  based  on  microscope  comparison
with  type  material  stored  in  the  NHMW  collection.  Except
for the about 2.5  smaller size the specimen coincides very
well with the type specimens.

The species occurs throughout the Late Oligocene and

Miocene in the Central Paratethys, the Mediterranean Sea
and the Eastern Atlantic (Tejkal et al. 1967; Báldi 1973;
Steininger 1973; Schultz 2001).

Genus: Leionucula Quenstedt, 1930

Leionucula ehrlichi

(Hoernes, 1875) nov. comb.

Fig. 9.2

1875 Nucula Ehrlichi nov. sp. – Hoernes, p. 378—379, pl. 14, figs. 11—13
1954 Nucula ehrlichi R. Hoernes – Csepreghy-Meznerics, p. 62, 121,

pl. 13, fig. 14

2001 Nucula (?Nucula) ehrlichi Hoernes, 1875 – Schultz, p. 1—2, pl. 1,

figs. 1—2

Fig. 8. 1—2 – Gadilina taurogracilis Sacco, 1897, 3—5 – Gadila gracilina Sacco, 1897, 6—7 – Fissidentalium badense (Partsch in
Hörnes, 1856).

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M a t e r i a l: 1 articulated shell embedded in sediment with

left valve exterior up, 1 right valve with peeled off exterior layer
(samples: 13—15, 17 and 18); length: 8.5 mm, height: 6.5 mm.

R e m a r k s: Shell small, low convex, longer than high with

ellipsoid outline, umbo slightly pointed and shifted posterior-
ly, shell wall thin, outer layer dull, inner layer shinny, shell ex-
terior ornamented by very fine growth lamellae, interior
margin not crenulated.

Except for the about two times smaller size, the specimen

corresponds fully to the type specimens from Ottnang in their
oval  outline,  thin  shell  wall  and  low-convex  valve.  The  ab-
sence  of  a  crenulation  of  the  interior  shell  margin  was  not
mentioned by Hoernes (1875) but can be seen in the original
illustrations and the reproductions by Schultz (2001) and in all
specimens  from  the  type  locality  housed  in  the  NHMW  col-
lection. In consequence of the latter feature, the species name
is now combined with Leionucula. The difference to L. laevi-
gata (Sowerby, 1818) is the more symmetrical outline result-
ing from the less posteriorly shifted umbo. The latter Neogene
species and its Paleogene predecessor (or synonym) L. pereg-
rina (Deshayes, 1858) have a reduced posterior shell portion
and  develops  a  stronger  postero-anterior  shell  elongation.
Based  on  its  outline,  the  shells  described  as  N.  laevigata  by
Tejkal et al. (1967) represent also Leionucula ehrlichi.

The species is restricted to the Ottnangian to Badenian of

the Central Paratethys, found between the Alpine Foredeep
and the North-Hungarian Basin (Tejkal et al. 1967; Steininger
1973; Schultz 2001).

Order: Nuculanoida Adams & Adams, 1858

Superfamily: Nuculanoidea Adams & Adams, 1858

Family: Yoldiidae Habe, 1977

Genus: Yoldia Möller, 1842

Yoldia nitida

(Brocchi, 1814)

Fig. 9.3—8

1814 Arca nitida: nob. – Brocchi, p. 482—483, pl. 11, fig. 3
1865 Leda nitida Brocc. – Hörnes, p. 308—309, pl. 38, fig. 9

1898 Yoldia nitida (BR.) – Sacco, p. 57—58, pl. 12, figs. 14—17
1989 Yoldia (Yoldia) nitida (Brocchi, 1814) – Andres, p. 328—329, pl. 1,

fig. 12

2001 Yoldia (Yoldia) nitida (Brocchi, 1814) – Schultz, p. 26, pl. 2,

figs. 3—4 (cum syn.)

M a t e r i a l: 2 articulated specimens, 8 left and 10 right

valves (samples: 5—6, 12, 16—17, 17, 17—18, 18—19, 19, 19—20,
20, 21—22 and scree material); LV – length: 11.8 mm, height:
6.8 mm, convexity: 1.8 mm.

R e m a r k s: Shell small, thin walled, moderately convex,

dorsally inflated. Dorsal margin trigonal with pointed umbo
projecting  over  the  hinge;  ventral  margin  broadly  convex,
posterior  margin  narrowly  convex  pointed;  anterior  margin
with broad rostrum. Shell exterior smooth with dorsally and
anteriorly developed fine, projecting, concentric lirae. Inte-
rior  margin  smooth,  hinge  taxodont  with  numerous  fine
teeth. Some specimens tend to broaden the anterior margin
similar  to  Y.  longa  (Bellardi,  1875)  from  the  Lower  Mio-
cene of Ottnang in Lower Austria. The latter species is less
convex and smooth, missing the previously described con-
centric sculpture.

This species, originally described from the Upper Pliocene

of  Northern  Italy,  is  represented  throughout  the  Miocene,
Pliocene and Pleistocene of the Paratethys, Mediterranean and
E  Atlantic.  In  the  Paratethys  the  oldest  well  documented
record  is  from  the  Ottnangian  of  the  North  Alpine  Foreland
Basin (Schultz 2001).

Order: Solemyoida Dall, 1889

Suborder: Solemyina Dall, 1889

Superfamily: Solemyoidea Adams & Adams, 1857

Family: Solemyidae Gray, 1840

Genus: Solemya Lamarck, 1818

Solemya doderleini

Mayer, 1861

Fig. 10.1—2

1861 Solenomya Doderleini Mayer – Mayer, p. 364
1865 Solenomya Doderleini Mayer – Hörnes, p. 257, pl. 34, fig. 10

Fig. 9. 1 – Nucula mayeri (Hörnes, 1865), 2 – Leionucula ehrlichi (Hoernes, 1875), 3—8 – Yoldia nitida (Brocchi, 1814).

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1875 Solenomya Doderleini Mayer – Hoernes, p. 376, pl. 13, figs. 9—12
1901 Solenomya Doderleini (May.) – Sacco, p. 128—129, pl. 27, figs. 1—4
2001 Solemya doderleini (Mayer, 1861) – Schultz, p. 29—31, pl. 2, fig. 8

(cum syn.)

M a t e r i a l: 4 poorly preserved articulated specimens (sam-

ple: 16—17); length: 6 mm, height: 2.5 mm.

R e m a r k s: Small, fragile, dorso-ventrally elongated shells

with ellipsoid outline. Low shell convexity and ornamentation
by  flattened  radial  folds  and  grooves  on  the  exterior  surface
which  are  posteriorly  more  prominent  and  anteriorly  sup-
pressed.  The  specimens  are  extremely  small  and  may  repre-
sent  juveniles.  They  show  identical  orientations  and  are
embedded within one single rock slab.

The species is present throughout the Oligocene to Miocene

in deep marine water deposits of the Mediterranean Sea and
from the Kiscellian to Badenian in the Paratethys Sea (Löffler
1999; Schultz 2001).

Subclass: Autobranchiata Grobben, 1894

Superorder: Pteriomorphia Beurlen, 1944

Order: Pteriida Newell, 1965

Suborder: Pinnina Waller, 1978

Superfamily: Pinnoidea Leach, 1819

Family: Pinnidae Leach, 1819

Genus: Atrina Gray, 1842

Atrina pectinata

(Linnaeus, 1767)

Fig. 10.3

1767 Pinna pectinata Linnaeus, p. 1160, nr. 264
1867 Pinna Brocchi d’Orb. – Hörnes, p. 372—373, pl. 50, fig. 2
1914 Atrina Basteroti no sp. – Cossmann & Peyrot, p. 67—69, pl. 11,

figs. 31—32

2001 Atrina pectinata brocchii (d’Orbigny, 1852) vel nov. ssp. –

Schultz, p. 133—136, pl. 10, figs. 5—6, pl. 11, fig. 1 (cum syn.)

M a t e r i a l: 1 left valve fragment and 2 shell fragments

(sample 17 At.B). Length/height of left valve fragment
31.0 mm/50.7 mm.

R e m a r k s: Shell only fragmentarily preserved, with dull

surface layer mostly flaking off remaining shiny, nacreous in-
ner shell layer. It is flattened, elongated posteroanteriorly and
thin walled, bearing at its proximal exterior surface fine radial
ribblets. Its central and distal exterior bear coarse ribs dorsally
and coarse irregular comarginal rugae ventrally.

There is ongoing discussion of the subspecies level taxono-

my of A. pectinata (Pfister & Wegmüller 1994; Schultz 2001).
We  consider  “Pinna  Brocchi  d’Orb.”  in  any  case  for  a  mor-
photype of A. pectinata and follow the taxonomic concept of
Studencka  et  al.  (1998).  The  taxonomic  revision  by  Marquet
(1995)  distinguishing  between  P.  pectinata  and  P.  fragilis
(Pennant,  1777)  and  proposing  the  renewed  introduction  of
distinguishing fossil subspecies names is not followed by sub-
sequent  authors.  This  common  species  in  Eggenburgian  to
Badenian deposits of the Paratethys still lives in world oceans
and seas. It occurs from the tide level up to bathyal depths.

Order: Pectinoida Adams & Adams, 1858

Superfamily: Pectinoidea Rafinesque, 1815

Familiy: Propeamusiidae Abbott, 1954

Genus: Parvamussium Sacco, 1897

Parvamussium felsineum

(Foresti, 1893)

Fig. 11.1—4

1893 Amussium felsineum Foresti – Foresti, p. 381—382
1897 Variamussium felsineum (For.) – Sacco, p. 49, pl. 14, figs. 7—22
1928 Amussium felsineum Foresti – Depéret & Roman, p. 182—183,

text-fig. 5, pl. 27, figs. 7—12

1977 Propeamussium (Parvamussium) felsineum (Foresti, 1895) –

Jakubowski & Musiał, p. 91—92, pl. 5, figs. 11—13, pl. 6, figs. 1—2

2001 Propeamussium (Parvamussium) felsineum (Foresti, 1893) –

Schultz, p. 162, pl. 15, figs. 10—11

M a t e r i a l: 7 articulated shells, 6 single left valves and 7

single right valves (samples: 2 L + 2 A , 7: 1 A, 10: 1 A,
13—14:  1 R,  16—17:  1 A,  17:  1 R + 1 L,  17—18:
1 A+2 R+ 1 L,  18:  1 R,  19:  1 A+ 2 L+1 R,  21:  1 R;
other  samples  –  7—8,  8—9,  15—16,  19—20,  20—21  and  scree
material).  Dimensions:  LV  –  height:  10.5 mm,  length:
10.2 mm, convexity: 0.6 mm.

R e m a r k s: Shell as high as long, low convex, circular in

outline, inclined anteriorly, dorsal margin straightened with
posterior ear small and anterior ear large and pointed. Right
valve exterior with regular, fine, slightly projecting comargin-
al lirae, left valve exterior with numerous (about 30 at central
disc portion) fine ribblets intersected by fine projecting lirae;
the number of ribblets increases distally by intercalation; scaly
nodes present at lirae/ribblet intersections. Interior with about
12 interior ribs restricted to proximal four fifths of height, the
very distal fifth with thinned shell wall.

Fig. 10. 1—2 – Solemya doderleini Mayer, 1861, 3 – Atrina pectinata (Linnaeus, 1767).

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The presence of radial ribblets on the left valve exterior sur-

face allows a differentiation of this species from P. duodecim-
lamellatum.  In  contrast  to  NW  Bulgaria  (Kojumdgieva  &
Strachimirov 1960), no smooth left valves were recorded from
the  Styrian  and  Vienna  Basins  and  the  identifications  of  P.
duodecimlamellatum in those basins is questionable (Deperet
&  Roman  1928;  Mikuž  1998;  Schultz  2001).  Because  of  the
prominent left valve sculpture, our specimens correspond to P.
felsineum morph. styriaca (Meznerics 1936).

P. felsineum is frequent in the Early Badenian of the Styrian

Basin  and  the  northern  Slovenia  (Meznerics  1936;  Mikuž
1998) and in the Early/Middle Badenian of the Vienna Basin
in  Austria  (Depéret  &  Roman  1928;  Schultz  2001)  and  NW
Bulgaria, the Late Badenian of the Carpathian Foredeep in SE
Poland (Jakubowski & Musiał 1977) and the Pannonian Basin
in  NE  Hungary  (Csepreghy-Meznerics  1960,  1966).  Further,
the species is present from the Middle Miocene to Pliocene in
the Mediterranean Sea and the NE Atlantic. The type locality
of Ponticello in the Val de Savena near Bologna in Italy is of
Pliocene age.

Order: Limida Waller, 1978

Superfamily: Limoidea Rafinesque, 1815

Family: Limidae Rafinesque, 1815

Genus: Limea Bronn, 1831

Limea strigilata

(Brocchi, 1814)

Fig. 12.1—4

1814 Ostrea strigilata – Brocchi, p. 571, pl. 14, fig. 15
1867 Lima strigilata Brocc. – Hörnes, p. 392, pl. 54, fig. 7
1898 Limea strigilata (Br.) – Sacco, p. 21—22, pl. 6, figs. 4—7
1907 Limea strigilata Br. sp. – Cerulli-Irelli, p. 89, pl. 4, fig. 45
1914 Limea strigillata mut. subhelvetica nov. mut. – Cossmann &

Peyrot, p. 160—161, pl. 20, figs. 37—38

1984 Limea (Limea) aff. strigilata (Brocchi, 1814) – Janssen, p. 57,

pl. 26, fig. 4

M a t e r i a l: 1 articulated specimen, 5 left and 2 right valves

(samples: 5—6, 7, 13, 16—17, 17—18, and scree material). Di-
mensions: LV – height: 6.8 mm, length: 5.7 mm, convexi-
ty: 1.8 mm.

Fig. 11. 1—4 – Parvamussium felsineum (Foresti, 1893).

Fig. 12. 1—4 – Limea strigilata (Brocchi, 1814), 5—7 – Limaria labani (Meznerics, 1936).

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R e m a r k s: Shell very small, highly inflated, anteroventral-

ly  elongated  with  straightened  hinge  margin  and  umbo  pro-
jecting over it. Very proximal and dorsal lateral shell portion
except  for  comarginal  lirae  smooth.  The  rest  of  exterior  sur-
face  bears  numerous  fine,  flat  topped,  smooth  radial  ribblets
intercalated at the distal margin by single scaly ribblets. The
whole  exterior  surface  bears  weak  comarginal  lirae,  better
visible in the rib interspaces and lateral dorsal margins. Interior
shell margin is crenated.

This species differs from Limaria tuberculata by the much

smaller,  not  gaping  shell,  showing  a  crenulated  interior  mar-
gin. The Middle Miocene L. subhelvetica from the Aquitaine
Basin  represents  a  somewhat  stronger  elongated,  less  convex
morphotype of L. strigilata. In the Paratethys, Limea strigilata
has  been  previously  described  only  from  Badenian  strata.
Therefore, the specimens from Cerová represent the stratigraph-
ically oldest occurrence of this Miocene to Pliocene species.

Genus: Limaria Link, 1807

Limaria labani

(Meznerics, 1936)

Fig. 12.5—7

1936 Lima (Mantellina) lábáni nov. spec. – Meznerics, p. 127, 133,

pl. 4, figs. 9—14

1967 Lima (Mantellum) labani Meznerics, 1935 – Tejkal et al., p. 162,

pl. 1B, fig. 22

2001 Limaria (Mantellina) labani (Meznerics, 1936) – Schultz, p. 301

M a t e r i a l: 2 left and 2 right single valves (samples: 18 and

21, and scree material); length: 8.5 mm, height: 6 mm.

R e m a r k s: Shell up to 17 mm long, thin walled, fragile, el-

lipsoid  in  outline,  strongly  elongated  postero-ventrally,  low
convex.  Hinge  line  short,  slightly  overwhelmed  by  umbo,
which is slightly inflated and pointed dorsally. Exterior sculp-
ture comprises coarse, regular comarginal folds and very fine
radial  ribblets  absent  only  at  dorsal  marginal  area  which  is
smooth except for fine growth lines.

Limaria labani is restricted to Karpatian and Badenian of

the Paratethys, occurring in the Slovenian part of the Styrian
Basin (Lower Badenian), the northern margin of the Pannon-
ian Basin in N Hungary (Karpatian—Badenian; Csepreghy-
Meznerics 1954) and E Slovakia (Karpatian) and in the
Vienna Basin in N Slovakia (Karpatian).

Subclass: Heterodonta Neumayr, 1884

Order: Venerida Adams & Adams, 1856

Superfamily: Tellinoidea Blainville, 1814

Family: Tellinidae Blainville, 1814

Subfamily: Macominae Olsson, 1961

Genus: Macoma Leach, 1819

Macoma elliptica

(Brocchi, 1814)

Fig. 13.1—2

1814 Tellina elliptica, p. nob. – Brocchi, p. 513, pl. 12, fig. 7
1875 Tellina ottnangensis nov. sp. – Hoernes, p. 370—371, pl. 13,

figs. 1—4

1879 Tellina Floriana. – Hilber, p. 418, 450, 451, pl. 6, figs. 1—2
1900 Tellina Floriana Hilber – Bauer, p. 38, 42—43, pl. 2, fig. 13
1900 Tellina Floriana var. plicata Bauer – Bauer, p. 43, pl. 2, fig. 14
1901 Macomopsis elliptica (Br.) – Sacco, p. 107—108, pl. 22, figs.

36—40

1910 Macoma elliptica (Brocchi) – Cossmann & Peyrot, p. 281—282,

pl. 9, figs. 33—35

1998 Macoma elliptica (Brocchi, 1814) – Duckheim & Strauch,

p. 203—219, 224—225, pl. 1, figs. 1—8, pl. 2, figs. 1—8

2001 Macoma (Psammacoma) elliptica (Brocchi, 1814) – Harzhauser

& Mandic, p. 748—749, pl. 9, fig. 8

2005 Macoma (Psammacoma) elliptica (Brocchi, 1814) s.l. (excl. M.

(P.) elliptica ottnangensis (Hoernes, 1875)) – Schultz, p. 721,
pl. 99, fig. 6 (cum syn.)

2005 Macoma (Psammacoma) elliptica ottnangensis (Hoernes, 1875)

– Schultz, p. 722—724, pl. 99, figs. 7—9

2005 Macoma (Psammacoma) elliptica floriana (Hilber, 1879) –

Schultz, p. 724—725, pl. 99, figs. 10—13

2005 Macoma (Psammacoma) elliptica plicata (Bauer, 1900) –

Schultz, p. 725, text-fig. 14

Fig. 13. 1—2 – Macoma elliptica (Brocchi, 1814), 3—4 – Lucina callipteryx Tournouer, 1874, 5—6 – Gonimyrtea submichelottii (Sacco,
1901), 7 – Thyasira flexuosa (Montagu, 1803), 8 – Laternula fuchsi (Hoernes, 1875).

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M a t e r i a l: 2 articulated specimens, 3 left and 5 right

valves (sample: 12, 13, 16, 16—17, 17—18, 19, 21+); LV –
length: 15.2 mm, height: 10.6 mm, convexity: 2.1 mm.

R e m a r k s: Shell small, thin walled, with shiny surface,

moderately convex, beak slightly pointed, shifted more or less
posteriorly,  dorsal  margin  trigonal,  ventral  margin  broadly
convex,  anterior  margin  narrowly  convex,  posterior  margin
pointed,  shell  exterior  smooth  with  minute,  weak  irregular
growth lamellae getting little more prominent only at posterior
margin,  interior  shell  with  smooth  margin,  deep  sinus  and
minute radial threads.

As pointed out already by Hoernes (1875) and Hilber

(1879)  this  is  a  highly  variable  species  especially  in  outline
and  convexity.  For  different  morphotypes  numerous  species
and  subspecies  names  are  available  in  the  literature.  In  the
present study, we follow the revision by Duckheim & Strauch
(1998) and consider those names as synonyms of M. elliptica.
Our  specimens,  with  more  centrally  placed  umbonal  region
and reduced posterio-ventral elongation, resembles particular-
ly Tellina floriana Hilber, 1879. Gastrana fragilis, in contrast,
may develop a similar outline but has a more prominent con-
centric sculpture showing well projecting concentric lirae.

The species is present from the Late Oligocene to Pliocene

in the Paratethys Sea, the Mediterranean Sea and the Eastern
Atlantic (Schultz 2005).

Superfamily: Lucinoidea Fleming, 1828

Family: Lucinidae Fleming, 1828

Subfamily: Lucininae Fleming, 1828

Genus: Lucina Brugui

re, 1797

Lucina callipteryx

Tournouer, 1874

Fig. 13.3—4

1874 Lucina callipteryx – Tournouer, p. 306, pl. 10, fig. 4
1911 Miltha (Eomiltha) callipteryx (Tournouer) – Cossmann & Peyrot,

p. 285—287, pl. 27, figs. 18—21

M a t e r i a l: 4 articulated specimens, 4 single left and 2 sin-

gle right valves (samples: 5—6, 7, 8, 13—15, 16, 16—17, 19,
19—20, 21). Dimensions: RV – length: 22.8 mm, height:
17.4 mm, convexity: 1.0 mm.

R e m a r k s: Shell small in size, low convex with postero-

dorsal depression; outline subquadrangular with subtrigonal
dorsal portion and rounded ventral portion; anterior margin
pointed, posterior margin rounded. Exterior surface with pro-
jecting comarginal lirae (0.5 mm distance in proximal part and
2.3 mm in distal part). Numerous fine growth lines are visible
between the lirae. Interior margin smooth.

The type specimen differs only in its about two times larger

size. Our specimens differ from Lucinoma borealis (Linnaeus,
1767) not only by more regular and more widely spaced con-
centric lamellae but mainly by the subquadrangular, anteriorly
pointed outline. The specimen from the Karpatian schlier of the
North Hungarian Basin, illustrated by Csepreghy-Meznerics
(1954) as Miltha ottnangensis (Hoernes, 1875), has a fragment-
ed anterior outline but the rest of the outline together with shell
exterior sculpture suggests an affinity to L. callipteryx.

This extraordinarily rare species, originally defined from the

Burdigalian of the Aquitanian Basin, was previously recorded
in the Central Paratethys only from the Lower Badenian of
Romania (Studencka et al. 1998). The present study provides
the first illustrations of a specimen ever found in the Para-
tethys Sea area.

Subfamily: Myrteinae Chavan, 1969

Genus: Gonimyrtea Marwick, 1929

Gonimyrtea submichelottii

(Sacco, 1901) nov. comb.

Fig. 13.5—6

1901 Dentilucina Meneghini de Stef. var. submichelottii – Sacco, p. 85,

pl. 20, figs. 10—11

1934 Phacoides submichelottii Sacco – Friedberg, p. 107, pl. 19,

figs. 4—5

M a t e r i a l: 10 articulated specimens and 1 right and 2 left

valves (samples: 7, 8—9, 12, 15—16, 16, 16—17, 17—18, 19,
19—20, and scree material). Dimensions: LV – length:
7.0 mm, height: 6.8 mm, convexity: 0.8 mm.

R e m a r k s: Shell is small, moderately convex with mod-

erately thick wall; outline rounded, exterior surface with
prominently projecting lamellae throughout ontogeny, some-
what narrower proximally than distally, interior shell margin
smooth.

Studencka et al. (1998) place this species into Parvilucina

which has – in contrast to Gonimyrtea – a crenated interior
shell  margin.  Phacoides  sub-Michelottii  (Sacco,  1901)  of
Cossmann  &  Peyrot  (1912,  pl. 28,  figs. 51—54)  from  the
Aquitanian Basin has a much finer concentric sculpture and a
stronger  developed  anterior  muscle  scar.  Therefore,  we
consider this identification as questionable. The illustrated
specimen  might  rather  represent  Gonimyrtea  meneghinii
(De  Stefani  &  Pantanelli,  1878)  from  the  Italian  Mio-
Pliocene that occurs in the Badenian of the Paratethys (Schultz
2001) as well. The very similar Lucina ottnangensis (Hoernes,
1875) is up to 4 times larger. It has a variably prominent con-
centric sculpture and is actually treated as Lucinoma borealis
(see Schultz 2003).

Gonimyrtea submichelottii is common in the Lower to Mid-

dle Miocene of the Turin Hills and in the Italian Pliocene. In
the Paratethys it was previously found only in the Lower
Badenian of southern Poland.

Family: Thyasiridae Dall, 1901

Genus: Thyasira Leach in Lamarck, 1818

Thyasira flexuosa

(Montagu, 1803)

Fig. 13.7

1803 Tellina flexuosa – Montagu, p. 72—73
1984 Thyasira (Thyasira) flexuosa (Montagu, 1803) – Janssen, p. 60—61,

pl. 1, figs. 4—5

1986 Thyasira (Thyasira) flexuosa (Montagu, 1803) – Studencka,

p. 56—57, pl. 7, fig. 10

2005 Thyasira (Thyasira) flexuosa (Montagu, 1803) – Marquet, p. 8—9,

pl. 2, fig. 2

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M a t e r i a l: 1 left valve (sample: 16—17); length: 6.0 mm,

height: 4.4 mm.

R e m a r k s: Minute shell, thin walled, moderately convex

with strong dorsal anterior depression, beak prosogyr, outline
subquadrangular rounded, exterior surface smooth with irreg-
ular, non-prominent concentric ribs.

The species is present from the Late Oligocene in the Para-

tethys, the Mediterranean and the NE Atlantic. In the Central
Paratethys its previous record is restricted to the Badenian
(Studencka et al. 1998), with one single specimen collected so
far from the Lower Badenian of Poland (Studencka 1986).

Subclass: Anomalodesmata Dall, 1889

Order: Pholadomyida Newell, 1965

Suborder

Pholadomyina Newell, 1965

Superfamily: Thracioidea Stoliczka, 1870

Family: Laternulidae Hedley, 1918

Genus: Laternula Röding, 1798

Laternula fuchsi

(Hoernes, 1875)

Fig. 13.8

1875 Anatina Fuchsi nov. sp. – Hoernes, p. 366—367, 393, 397, pl. 13,

figs. 13—16

1954 Anatina fuchsi R. Hoernes – Csepreghy-Meznerics, p. 108, 126,

pl. 13, fig. 15

1973 Laternula fuchsi (R. Hoernes, 1875) – Steininger, p. 542, pl. 30, fig. 1
1998 Laternula (Laternula) fuchsi (Hoernes) – Schultz, p. 112, pl. 50,

fig. 5

2005 Laternula (Laternula) fuchsi (Hoernes, 1875) – Schultz,

p. 1043—1044, pl. 151, figs. 2—4

M a t e r i a l: 7 articulated specimens, and 2 right valves

(samples: 13, 13—14, 15, 15—16, 16, 16—17, 20—21). Dimen-
sions: LV – height: 12.3 mm, height: 10.6 mm, convexity:
1.6 mm.

R e m a r k s: The shell is small, thin walled, shiny and trans-

lucent,  low  convex  with  oblique  posteroventral  depression;
outline  oval  to  rounded,  elongated  anteroventrally,  anteriorly
rounded,  posteriorly  truncated,  umbo  posteriorly  pointed.
Shell exterior smooth, with fine irregularly projecting comar-
ginal lirae and/or rugae.

As already remarked by Steininger (1973), the shells of this

species are usually highly deformed and the outline is seem-
ingly  very  variable.  The  only  well  preserved  specimen
(Fig. 13.8)  displays  a  rather  rounded  outline  whereas  other
specimens  show  a  posteroventral  elongation  which  is  typical

for the type specimen from Ottnang in Upper Austria (NHMW
collection). Therefore, the specimens differ from other L.
fuchsi (Hoernes, 1875) morphs only by their about 2 to 3
times smaller size.

The previous fossil record of this species is restricted to the

Ottnangian of Upper Austria and the Karpatian to Lower Bad-
enian of the North Hungarian Basin (Csepreghy-Meznerics
1954). Additionally, it has been reported from the Ottnangian
of Bavaria (Steininger 1973), the Karpatian of Slovakia (Tej-
kal et al. 1967) and the Lower Badenian of Northern Slovenia
(Meznerics 1936).

Suborder: Cuspidariina Dall, 1886

Superfamily: Cuspidarioidea Dall, 1886

Family: Cuspidariidae Dall, 1886

Genus: Cardiomya Adams, 1864

Cardiomya elegantissima

(Hoernes, 1875)

Fig. 14.1—3

1875 Neaera elegantissima M. Hoernes – Hoernes: p. 368, pl. 13, fig. 8
1967 Cuspidaria (Cuspidaria) elegantissima (R. Hoernes, 1875) – Tej-

kal et al., p. 189, pl. 8B, fig. 16

1973 Cuspidaria (Cuspidaria) elegantissima (Hoernes, 1853) – Steinin-

ger, p. 544, pl. 30, fig. 2

2003 Cuspidaria (Cuspidaria) elegantissima (Hoernes, 1875) – Schultz,

p. 1056, pl. 152, fig. 3

M a t e r i a l: 5 articulated specimens, 2 left valves and 1 right

valve (sample numbers: 5—6, 6—7, 7—8, 13—14, 16, 17—18,
19—20 and scree material). Dimensions: length: 10.5 mm,
height: 6 mm.

R e m a r k s: Shell small, about 10 mm in length, fragile,

postero-ventrally elongated, and moderately convex. Anterior
side  slightly  pointed,  narrowly  rounded,  with  shell  surface
bearing coarse irregular concentric ribs. Posterior side shortly
rostrate,  with  posterior  part  of  the  exterior  surface  bearing
three prominent radial triangular ribs, laterally adjoined by up
to  four  additional  tinny  ribs;  rest  of  exterior  surface,  at  ros-
trum, smooth. This species is reminiscent of the extant C. cos-
tellata (Deshayes, 1833) from the Mediterranean Sea and the
Atlantic, which is a carnivorous deep water bivalve (Poutiers
& Bernard 1995). The latter differs from C. elegantissima by
much weaker concentric ribs at the anterior shell surfaces.

This species is restricted to the Lower Ottnangian of the

eastern North Alpine Foredeep and the Karpatian of the north-
ern Vienna Basin.

Fig. 14. 1—3 – Cardiomya elegantissima (Hoernes, 1875).

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Mollusc taxa from Cerová

Gastropoda

Calliotropis? sp.
Polinices cerovaensis nov. sp.
Stellaria testigera (Bronn, 1831)
Galeodea echinophora (Linnaeus, 1758)
Mitrella hilberi (Cossmann, 1901)
Nassarius janschloegli nov. sp.
Amalda glandiformis (Lamarck, 1810)
Conolithus antidiluvianus (Brugui

re, 1792)

Genota valeriae (Hoernes & Auinger, 1891)
Pyramidella cf. gratteloupi d’Orbigny, 1852
Turbonilla sp.
Triphora sp.
Ringicula minor (Grateloup, 1838)
Cylichna cf. salbriacensis (Peyrot, 1932)
Sabatia callifera Boettger, 1906
Balantium collina (Janssen & Zorn, 2001)

Scaphopoda

Fissidentalium badense (Partsch in Hörnes, 1856)
Gadilina taurogracilis Sacco, 1897
Gadila gracilina Sacco, 1897

Bivalvia

Nucula mayeri Hörnes, 1865
Leionucula ehrlichi (Hoernes, 1875)
Yoldia nitida (Brocchi, 1814)
Solemya doderleini Mayer, 1861
Atrina pectinata (Linnaeus, 1767)
Parvamussium felsineum (Foresti, 1893)
Limea strigilata (Brocchi, 1814)
Limaria labani (Meznerics, 1936)
Macoma elliptica (Brocchi, 1814)
Lucina callipteryx Tournouer, 1874
Gonimyrtea submichelottii (Sacco, 1901)
Thyasira flexuosa (Montagu, 1803)
Laternula fuchsi (Hoernes, 1875)
Cardiomya elegantissima (Hoernes, 1875)

Acknowledgments: We thank Daniela Esu (Dip. Scienze del-
la Terra, Universit

Sapienza, Roma) for her constructive re-

view, and to Natália Hudáčková, Andrej Ruman and Matúš
Hyžný (Comenius University in Bratislava) for their help dur-
ing the field and laboratory works. The work has been sup-
ported by research Grant APVV 0280-07 and by Grant KEGA
3/7226/09 and contributes to the FWF-Project P23492.

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