www.geologicacarpathica.sk
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
, APRIL 2011, 62, 2, 121—138 doi: 10.2478/v10096-011-0011-1
Introduction
The Krížna Unit in the eastern part of the Malá Fatra Mts is
formed by the Upper Triassic carbonate complexes, which
have been studied in the Ve ká Lučivná Valley between
Zázrivá and Párnica villages (Fig. 1). Dolomite formations in
the Ve ká Lučivná Valley cropped out along the forest roads,
where the dark grey, yellowish-grey and grey-green clay-
stones and dolomitic shales were also uncovered. Dolomite
beds and shaly interbeds occur in lens-shaped bodies, which
are underlain by the Ramsau Dolomite Formation and over-
lain by the Hauptdolomite Formation (Fig. 2). The claystones
contain the rare fossils, such as lingulid brachiopods and con-
chostracans, which mean that these sediments represent a
unique facies unknown from the Upper Triassic formations of
the Krížna Unit up till now. Besides claystones and carbonates
the Upper Triassic sediments in the eastern part of the Malá
Fatra Mts are also represented by the Lunz Beds, which occur
rarely in the form of discontinuous lenslike bodies within the
Upper Triassic dolomites. Both sedimentary formations of
claystones and sandstones were probably deposited during the
Carnian. They represent the intraplatform and periplatform
terrigenous facies, which were deposited in response to the de-
mise of carbonate platforms and enhanced siliciclastic input
during the Reingraben Event (Schlager & Schöllnberger
1974), Carnian productivity crisis (Hornung et al. 2007a),
Carnian Pluvial Event (Simms & Ruffel 1989) and Carnian
humid intermezzo (Kozur & Bachmann 2010).
Siliciclastics in the Upper Triassic dolomite formations of the
Krížna Unit (Malá Fatra Mountains, Western Carpathians):
constraints for the Carnian Pluvial Event in the Fatric Basin
MILAN SÝKORA
1
, MILOŠ SIBLÍK
2
and JÁN SOTÁK
1,3,4
1
Department of Geology and Paleontology, Faculty of Natural Sciences, Comenius University, Mlynská dolina G, 842 15 Bratislava,
Slovak Republic; sykora@fns.uniba.sk
2
Institute of Geology v.v.i., Academy of Sciences of the Czech Republic, Rozvojová 269, 165 00 Prague 6, Czech Republic; siblik@gli.cas.cz
3
Geological Institute, Slovak Academy of Sciences, Ďumbierska 1, 974 11 Banská Bystrica, Slovak Republic; sotak@savbb.sk
4
Department of Geography, Faculty of Education, Catholic University, Hrabkovská cesta 1, 03401 Ružomberok, Slovak Republic
(Manuscript received July 9, 2009; accepted in revised form December 16, 2010)
Abstract: The Upper Triassic carbonates of the Krížna Unit in the eastern part of the Krivánska Malá Fatra Mts contain
the unique siliciclastic interbeds. They are developed between the Ramsau Dolomite Formation and the Hauptdolomite
Formation, and comprise alternating dark grey shales and pale grey dolomites. Such a shaly-dolomite formation is
unknown from the Upper Triassic formations of the Krížna Unit, which makes it possible to define a new lithostratigraphic
unit (Tržinovo Formation). The Ramsau Dolomites beneath the Tržinovo Formation contain microfauna of Carnian
foraminifers. Therefore, the Carnian age has also been constrained for the Tržinovo Formation, which by the reduced
carbonate productivity and enhanced terrigenous influx could be related to the Reingraben Event. This event related to
the “Carnian Pluvial Episode” is also inferred in the Tržinovo Formation by the presence of lingulide brachiopods
(Lingularia) and spinicaudatan crustaceans (Euestheria). This fauna provides evidence of not fully marine conditions of
the Tržinovo Formation, influenced by continental freshwater influx and humid climate.
Key words: Carnian event, Upper Triassic dolomites, Western Carpathians, Krížna Unit, shaly interbeds, lingulids,
conchostracans.
Study area and geological setting
The Triassic sequence of the Krížna Unit in the area studied
is represented by limestone, dolomitic, sandy siliciclastic and
shaly claystone formations (see lithostratigraphical column in
Fig. 2).
The Middle Triassic sequence belongs to the Gutenstein
Limestone Formation (Anisian), composed of dark grey bed-
ded limestones with mudstone, wackestone and packstone mi-
crofacies, molluscs, echinoids, foraminifers and ostracods.
The overlying sequence of the Ramsau Dolomite Formation
(Anisian to Lower Carnian see Lexa et al. 2000 for compari-
son) consists of grey, bedded or massive, fine-grained to crys-
talline dolomites with rare bivalves, echinoids, peloids and
algae. This formation locally contains discontinuous lens-
shaped bodies of crinoidal and bioclastic limestones with
well-bedded stratification (indicated in Fig. 2). The Fassanian
age of these limestones has been determined by the conodonts
of Gondolella acuta Kozur and Gondolella trammeri Kozur
(Sýkora 2003).
The Ramsau Dolomite Formation, the uppermost part of
which belongs to the Carnian, is overlain by sediments, repre-
sented by two different clastic formations. The first is repre-
sented by the Lunz Beds, which are composed of fine-grained
sandstones (feldspathic graywackes mainly) with thin interca-
lations of sandy or silty shales. The carbonate-terrigenous fa-
cies is recognized herein as the Tržinovo Formation, which is
characterized by alteration of shales and dolomites. Both ter-
122
SÝKORA, SIBLÍK and SOTÁK
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
rigenous formations occur in lens-shaped bodies within the
Upper Triassic carbonate formations.
The terrigenous facies of the Lunz Beds and Tržinovo Forma-
tion was subsequently replaced by sedimentation of carbonates.
The Late Carnian and Norian Hauptdolomit Formation consists
of light grey bedded dolomites with very scarce allochems. The
overlying Norian Carpathian Keuper Formation is composed of
variegated shales, siltstones, quartzose sandstones and fine-
grained dolomites (so-called Keuper dolomites).
The uppermost formation of the Triassic sequence is repre-
sented by shallow-marine sediments of the Rhaetian Fatra
Formation, which consists of bedded limestones with cyclic
development of bioclastic, ooidal, oncoidal, peloidal and mi-
crobial limestones (lagoonal and patch reefs), interlayered
with shaly claystone and occasional dolomite beds.
The information about the presence of the siliciclastic inter-
beds in the Carnian formation of the Krížna Unit unknown un-
til the paper by Kozur & Mock (1993), who mentioned
occurrences and findings of fossils in the claystones from a
forest road outcrop in the Lučivná Hill area (see Fig. 1). A
similar formation of dolomitic clays and shaly clays has been
described from the Krížna Unit in the Ružbachy Horst by
Kullmanová & Nemčok (1985), where its Carnian age is
proved by lamellibranchs and foraminifers (see below). In-
creased terrigenous material in the Upper Triassic dolomites
underlying the Carpathian Keuper Formation is also recorded
in the Krížna Unit in some mountains of the Western Car-
pathians, like Považský Inovec Mts, Strážovské vrchy Mts,
Low Tatra Mts and Tatra Mts (see Mahe et al. 1967; Mahe
1986), but their more precise stratigraphic position and litho-
logical descrition remains unknown.
The carbonates and claystones of the Tržinovo Formation
have been studied in two sections designated as P—A locality
and P—B locality with a distance of about 1.5 km (Fig. 1). The
sections show the alternation of claystones and dolomites,
which resemble the sediments of the platform and shelf facies
of the Raibl Shales in the Norther Calcareous Alps (see e.g.
Tollmann 1976), but without the marine fossils typical for the
Raibl Shales and especially the Reingraben Shales in the Al-
pine and Western Carpathian units (Choč Unit). The fossil
components of the Tržinovo Formation (P—A locality) com-
prise only the infaunal brachiopods of the genus Lingula to-
gether with spinicaudatan crustaceans—conchostracans of the
family Euestheridae (Kozur & Mock l993).
The stratigraphic position of the Tržinovo Formation is in-
ferred from foraminifers in dolomitic limestones of the Ram-
sau Dolomite Formation. In the section studied here it directly
underlies the claystones with conchostracan species of Eues-
theria minuta (von Zieten), which according to Kozur &
Weems (2010) formed the Late Ladinian zone, but is still
present in the entire Cordevolian. The presence of these fresh-
wather to brackich-water conchostrachans in the Tržinovo
Formation reflects the Carnian carbonate productivity crisis in
the North Tethyan realm, which corresponds to the Reingra-
ben Event during the Early Carnian.
Exposure of the Lunz Beds occurs in the dolomites near the
Lučivná elevation point on the left (southern) slope of the
Tržinovo Valley. Its verified thickness is just about 10 m, and
its observable lenght is a few hundred meters (see Fig. 1).
Outcrops and debris derived from the Lunz Beds are situated
around 200 m to the NW from the P—A locality. The lithology
of the Lunz Beds is different from the shaly-carbonate sedi-
Fig. 1. Location and geological map of the surroundings of the Tržinovo Valley area with localities studied.
123
SILICICLASTICS IN TRIASSIC DOLOMITE: CONSTRAINTS FOR CARNIAN PLUVIAL EVENT (W CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
Macrofossils were collected from the claystones, and prepared
for paleontological study. The claystone samples from the P—
A-section (layers I, II, III see Fig. 3) were macerated and treat-
ed with HCl and HF by using of standard palynological
processing techniques, but preservation of the extracted pa-
lynomorphs was not sufficient for specific determination.
Several samples of the dolomitic limestones were dissolved
in 10% acetic acid to extract stratigraphically important mi-
crofossils, like conodonts. The carbonate (dolomitic) fraction
of the samples from the selected claystones (layers I, II, III,
IV) was analysed by manometric methods (Turanová & Turan
1993), and was removed by diluted HCl. The mineralogical
composition of the insoluble residue (fraction under 2 µm)
was studied by using X-ray powder analysis (PXRD). The
analysis was performed on a DRON-3 diffractometer operat-
ing at 40 kV and 15 mA, with CoK radiation at scan speed of
0, 02 °2 in range of 4—74 °2 . Oriented specimens were pre-
pared by the sedimentation from suspension on a glass slide.
The prepared samples were analysed at room temperatures
(air-dry) and after treating with ethylenglycol (EG) for 8 hours
at a temperature of 60 °C.
Thin sections of the samples are deposited in the Depart-
ment of Geology and Paleontology of Comenius University
Bratislava (M. Sýkora) and in the Geological Institute of the
Slovak Academy of Sciences Banská Bystrica (J. Soták). The
macrofossils Lingularia and Euestheria figured in this paper
are deposited in the Slovak National Museum in Bratislava.
Microfacies and petrographic analysis of the
sections
Section A (P—A)
The sequence of the P—A section can be divided into two
parts (see Fig. 3). The lower part of the section represents the
uppermost members of the Ramsau Dolomit Formation and
consists of dolomites and dolomitic limestones (beds No. 04,
03, 02, 01, 1, 1a, 1b, 2, 2a, 2b, 3, 4, 5b, 5v, 6, 7, 8, 8a). The
stratigraphic age of the Ramsau Dolomite Formation in the
Krížna Unit corresponds to Anisian up to Carnian—Cordevo-
lian (see Biely et al. 1997; Lexa et al. 2000). The Carnian age
is also indicated by the foraminiferal microfauna (see strati-
graphical study) from the terminal beds of the Ramsau dolo-
mite (beds No. 0 to 3). The lower part of the section is only
partly exposed with an observable thickness of about 6.5 m.
Four bedset groups of grey, partly dolomitized limestones, al-
ternating with grey to light grey dolomites can be distin-
guished here.
In the lower part of the section A (No. 04 to 03) the bedset
of dolomitic limestones are represented by packstones and
grainstones mainly with intraclasts, peloids and bioclasts.
Rounded intraclasts of mudstones, wackestones and peloids
are dominant. Foraminiferal components dominated by the
uniserial morphotypes, like Nodosaria and Pseudonodosaria.
Ostracods and shell fragments of bivalves are less abundant.
Algal fragments (indeterminable thallii of dasyclads) and
miliolid foraminifers Agathammina austroalpina Kristan-
Tollmann & Tollmann are very rare.
Fig. 2. The lithostratigraphic column of the Triassic sequences of
the Krížna Unit in the eastern part of the Malá Fatra Mts, indicating
the position of the Tržinovo Formation.
ments at P—A and P—B sections. In the Lunz Beds, the sand-
stones do not alternate with dolomites, but they are rather ho-
mogeneous and do not contain any claystones typical of the
sections studied. Thin lenslike bodies of the Lunz Beds s.l. are
also known from other areas of the Malá Fatra Mts (Haško &
Polák 1979; Rakús et al. 1988).
Material and methods
Sedimentary sequences have been logged, analysed and
sampled. Selected samples of carbonates were studied by mi-
croscopic methods to determine microfacies, petrographic fea-
tures and microfossils like foraminifers, algae, ostracods, etc.
124
SÝKORA, SIBLÍK and SOTÁK
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
on the basal bed surfaces. The limestones contain
flat mudstone lithoclasts with fine parallel lami-
nations and clasts of cryptalgal bindstones with
micritic algal biolamination. Peloids and small
biodetritic grains are also present.
Bed No. 1 is formed by foraminiferal grain-
stones with involutinid, ammodiscid, miliolid
and rarely duostominid foraminifers (Fig. 6b,c).
Dasyclad algae Halycorine and echinoderm frag-
ments were rarely found, as well. In this lime-
stone, the specific components are represented by
aggregate grains (Fig. 6a,b) forming grapestone
to lumpy aggregates (cf. Tucker & Wright 1990),
which indicate the shallow marine environments
of the subtidal to intertidal zone. Mixing of bio-
clasts and intraclasts derived from various sourc-
es reflects the changes in depositional energy and
intensity of bottom erosion (rip-up intraclasts).
Disorganization of some tempestite-like beds
(Fig. 5b, No. 0, 1) indicates a storm activity, but
typical storm-related textures like lags and scour-
and-fill structures are not developed.
The layers No. 1a,b, 2, 3 are composed of
grainstones, less mudstones, wackestones and
packstones. The beds provide the structures of
fenestral fabrics, fine lamination, bioturbation
and rare desiccation cracks filled by breccias.
Low-diversity microfauna is typical of these sed-
iments, revealing an abundance of nodosariid
foraminifers in some beds.
The superposed beds No. 4, 5 are formed by
dolomitic limestones with textures of mudstones,
packstones and grainstones. The special microfa-
cies fabric is recorded in the bed No. 4, which
consists of peloidal bioturbated wackestone with
irregular vuggs and tubular fenestrae (Fig. 6d).
These textures were probably produced from
trapping of gases released from the sediment.
The texture of bed No. 5 is heterogeneous. Its
basal part bears the lithoclasts of mudstones with
small irregular fenestrae (Fig. 5d), micritic intrac-
lasts and frequent peloids. Aggregate grains, intr-
aclasts and fenestral pores occur in the upper part
of this bed, associated with rare foraminifers, bi-
valves and ostracods. Laminoid fenestral fabrics
of the LF-BII-type have been observed, too. Ac-
cording to Flügel (2004), such fabrics are typical
of modern intertidal and supratidal environmets.
The described bedsets of dolomitic limestones
alternate with fine-grained dolomites. They con-
tain relatively frequent pseudomorphs of dolo-
mite after evaporitic minerals, such as sulphates.
Fig. 3. Lithological sections from the localities studied (Section A, B), extending from
the upper part of the Ramsau Dolomite Formation, through the shaly-dolomitic se-
quence of the Tržinovo Formation to the lower part of “Hauptdolomit” Formation.
In the upper bedsets (No. 02—3) the dolomicritic limestones
(mudstones, wackestones and packstones) alternated with
grainstones and breccias. The allochems are similar to the do-
lomitic limestones of the former bedset, apart from those in
the grainstone of bed No. 1 (foraminiferal limestones). Beds
No. 02, 01, 0 consist of the carbonatic breccias (Fig. 5b),
formed by increasing water energy with erosional wash-outs
Fossils (ostracods and uniserial foraminifers) are generally un-
common. In thin sections of dolomites, the admixture of silty
and very fine sandy quartz has also been identified in a few
cases (beds No. 6, 7, 8a).
The topmost beds of the Ramsau Formation (No. 8) are
composed of wackestone/floatstone limestones with micritic
intraclasts up to 7 mm in size (Fig. 5c). The micritic matrix
125
SILICICLASTICS IN TRIASSIC DOLOMITE: CONSTRAINTS FOR CARNIAN PLUVIAL EVENT (W CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
Fig. 4. Sedimentary sequence and internal structures of the Tržinovo Formation. a – Lower part of section A (P—A), locus typicus of the Tržinovo
Formation, formed by two members: Tržinovo Shales and Tržinovo Dolomites. Outcrop in forest road cut, hammer as a scale with 50 cm length.
b – Lower part of section B (P—B), parastratotype of the Tržinovo Formation, formed by two members. Dolomite layer exhibits a seismotectonic
deformation on the top of the claystone bed-marked by the arrow in the middle of the picture. Forest road cut outcrop, hammer as a scale. c – In-
traformational carbonate conglomerate deposited probably as a storm lag. Dark lithoclasts in lower part show the liesegang bands. Sample from
the Ramsau Dolomite Formation outcrop in the forest road cut near section A. d – Lithoclast with liesegang bands from carbonate conglomerate,
see previous picture.
also contains fine-sized biodetrital grains (less than 0.1 mm),
the orgin of which is uncertain. This type of micritic limestone
with well-rounded intraclasts prove evidence of textural inver-
sion, which is typical of the high energy environments of
channels. The lithology of the lower part of the section chang-
es above bed No. 8a, where the carbonates are suddenly re-
placed by dolomitic claystones, exhibiting no visible erosion
contact.
Samples of the dolomitic limestones from the beds No. 04
and 1 were dissolved in 10% CH
3
COOH, but no stratigraphi-
cally important microfossils were found in the insoluble
residuum.
The superposed formation in the upper part of section A is
formed by dolomitic claystones and claystones (I to VI beds),
intercalated by fine-grained, pale grey carbonates in the
beds 8b,c, 9. The sequence with alternation of shales and car-
bonates forms the upper part of the section (11.8 m), provid-
ing the various thickness of claystones in the lower bed I (up
to 120 cm) and uppermost bed VI (only 2 cm). The colour of
the claystones is grey, dark grey, greenish-grey and yelowish.
The carbonates are mostly fine-grained, pale grey dolomites
with specific allochems and detritic dolomitized limestones.
Bed No. 8b represents moderately to poorly sorted grainstone
to packstone with intraclasts, peloids, aggregate grains and fe-
nestrate cavities. Rare bioclasts have their origin in bivalves,
foraminifers, ostracods and gastropods. Bed No. 8c is formed
by breccias with flat clasts of laminated mudstones or bind-
stones; the second part of this bed consists of grainstone with
mudstone lithoclasts. Bed No. 9 is formed by breccia with
fine-grained matrix and poorly-sorted mudstone intraclasts,
which reveals the textural inversion. Therefore beds No. 8b,c
and 9 are considered to represent channel deposits. Allochems
of these carbonates have qualitatively similar composition to
the dolomitic limestones in the lower part of section A (beds
No. 04—5). Dolomites commonly have parallel lamination and
form generally thin-bedded layers.
The claystones are composed of illite and dolomite (XRD
and DTA analyses). The dolomite content in claystones in
the middle part of shaly beds is as follows: bed I – 34.4 %;
II – 7.2 %; III – 0 %; IV – 20.3 %. The shales are pelitic,
with the average grain size of 0.007 mm.
The dolomitic claystones in bed No. I are dark grey to grey
in colour. This bed contains the brachiopods and spinicauda-
tans (conchostracans), which was firstly recorded by Kozur &
126
SÝKORA, SIBLÍK and SOTÁK
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
Fig. 5. Microfacial types of the carbonates from the Raumsau Dolomite Formation. a – Dolomitic limestone with fine parallel lamination,
bed No. 01, thin section 27284, section A, scale 1 mm. Ramsau Dolomite Formation. b – Carbonatic microbreccia with mainly angular li-
thoclasts of fine-grained carbonates. Channel-type sediment, bed No. 0, thin section 27287, Ramsau Dolomite Formation, section A, scale
1 mm. c – Intraclasts of micritic carbonates with wackestone to packstone matrix, textural inversion in bed No. 8, thin section 27103,
Ramsau Dolomite Formation, section A. Scale bar 1 mm. d – Fenestral fabric with irregular voids in carbonates from lower part of bed
No. 5, thin section 27282a, Ramsau Dolomite Formation, section A, scale bar 1 mm.
Mock (1993). These fossils are relatively rare, and they occur
as isolated valves often with fragmentary preservation. The
claystones with lingulids do not show any burrowings and
presence of ichnofossils. The lingulids are arranged mostly
parallel to bedding, indicating a post-mortem transport of their
shells. Brachiopods of Lingularia ex gr. tenuissima (Bronn)
(description in systematic part) occurs much more frequently
than conchostracans of Euestheria minuta (von Zieten) (deter-
mination by Kozur & Weems 2010). Bioclasts were recorded
only in bed No. I. Other macrofossils have not been found, apart
from very rare small (less than 1 mm) fragments of fish? bones.
Besides fossils, the authigenic pyrite (in small grains below
0.1 mm) and rare oxidized pyrite concretions (max. size 2 cm)
were found in the claystones. The claystones occasionally
yield a very fine parallel lamination and admixture of silty ter-
rigenous quartz. The inorganic and organic carbon content of
the claystones is as follows: Bed No. I (dark grey claystones
with fossils) – TC 3.63 %, TOC 0.5 %, TIC 3.13 %; Bed
No. II (grey-brownish claystones without fossils) – TC
1.65 %, TOC 0.18 %, TIC 1.47 %; Bed No. III (grey-greenish
claystones without fossils) – TC 0.1 %, TOC trace, TIC
0.1 %; Bed No. IV (grey-yellowish dolomitic claystone –
5 cm thin) – TC 2.34 %, TOC 0.32 %, TIC 2.02 %. The clay
fractions of below 2 µm were determinated by XRD analysis
from the beds No. I, II and III. The dominant clay mineral was
illite. Kaolinite was determined only in the bed No. I (up to
5 %). The kaolinite/illite ratio is about 0.05, which indicates
humid and warm climate (compare to Simms & Ruffell 1989,
1990). Other clay minerals were not recorded. In the coarser
fraction of the claystones (over 2 µm), minerals like dolomite,
quartz and feldspar were identified.
Section B (P—B)
The thickness of the outcropping strata is about 11 m, the
sedimentary sequence is very similar to the formation de-
scribed from section A (see Fig. 4b). The lower part of the sec-
tion is built up by the Ramsau Dolomite Formation, formed by
grey, fine-grained dolomites and less frequently by dolomitic
limestones, just like those in section A. Bed No. 2 is com-
127
SILICICLASTICS IN TRIASSIC DOLOMITE: CONSTRAINTS FOR CARNIAN PLUVIAL EVENT (W CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
posed of grainstones with very similar microfacies and fora-
miniferal assemblage to those described from bed No. 1 in
section A. The dolomites occasionally contain pseudomorphs
after evaporites, cryptalgal fabrics and indistinctive parallel
lamination. Allochems are rare and occur mainly as bioclasts.
The texture and components of the limestones are very similar
to those of the lower part of section A.
The overlying claystones and dolomitic claystones corre-
spond to the claystones in the A-section by their composition
and colour. Similarly, they form 6 layers with various thick-
nesses, from 2 to 100 cm. The upper bedding plane of the
claystone bed I is dissected by short faults (see Fig. 4b
marked by arrow), which do not cut the superposed dolomite
layer. Overlying carbonate infill irregularities caused by the
synsedimentary faulting are probably due to seismic activity.
Therefore, it is supposed that these deformations were syn-
or slightly post-sedimentary. In the section A, the bedding
plane above the claystone I does not exhibit any deformation
structures. Neither macrofauna nor microfossils were found
in the claystones from section B.
Sedimentary environments
The Ramsau Dolomite Formation in the lower part of the
sections (A – 6 m, B – 2.8 m) represents peritidal carbon-
ates. The beds are grouped into four bedsets of dolomitic
limestones and dolomites (Fig. 3), representing high-fre-
quency cycles (thickness 1.4—1.7). Peritidal carbonates pro-
vide a shallowing-upward trend with appearance of parallel
lamination and fenestral fabrics (Figs. 5a,d, 6c,d). Intraclasts
are also frequent in the beds, especially in the lower and
middle part of cycles. They form rounded to angular clasts
with various sizes (Fig. 5b,c), consisting of mudstones,
wackestones and packstones with laminae of fine biodetrital
grains (below 0.1 mm). Supratidal conditions are reflected
by the presence of pseudomorphs of dolomite (after gypsum/
anhydrite) and peloids. Besides allochems, the grainstone
type limestones also contain aggregate grains—grapestones
(Fig. 6a,b,c). Some beds indicate the changes in depositional
energy, by which various sized grains were periodically
mixed and packed in micritic matrix.
Fig. 6. Microfacial types of the carbonates from the Ramsau Dolomite Formation. a – Bioclastic grainstone with allochems (foraminifers and
fragments of lamellibranchiate shells), peloids and aggregate grains of grapestone from the bed No. 1. Mixed grains and bioclasts implies a
poorly sorted tempestite. Scale 1 mm, thin section 27287b, section A. Ramsau Dolomite Formation, scale 0.5 mm. b – Bioclastic grainstone
with lamellibranchiate shells coated by sesile foraminifers and lumps of aggregate grains from bed No. 1. Scale is 1 mm, thin section
27287a, section A, Ramsau Dolomite Formation. c – Peloidal packstone, locally grainstone with foraminifers and laminoid fenestral fabrics,
bed No. 2a, thin section 27290, Ramsau Dolomite Formation, section A, scale 1 mm. d – Fenestral fabric with tubular fenestrae in wacke-
stone-type carbonates wackestone. The fenestral cavities were probably formed by degasation and escape of gas bubbles from soft sediment.
Bed No. 4, thin section 27281, section A, Ramsau Dolomite Formation, scale 1 mm.
128
SÝKORA, SIBLÍK and SOTÁK
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
Microfossils are represented mainly by foraminiferal as-
sociations from different shallow-water environments
(No. 02—1b and 2—2b in section A, No. 2 in section B). Aulo-
tortid- and involutinid-rich foraminiferal associations are
atypical for the Triassic carbonates of the Krížna Unit, where
they were found only for the second time. This type of fora-
miniferal microfauna is known almost exclusively from the
Hronic and Silicic Units.
Organic content and faunal diversity is low in general. Os-
tracods, fragments of bivalve shells, gastropods and green al-
gae are recorded in the thin sections. Some of the mollusc
shells are coated by thin micritic envelopes, due to activity
of epilithic organisms (see Kobluk & Risk 1977), cyanobacte-
ria and endolithic microborings. Epiplanktonic zoospores
(Globochaete) and shells of juvenile bivalves (“filaments”)
were identified very rarely. On the contrary, the coated grains,
grapestones, thick-walled foraminifers and fragments of dasy-
clad algae were derived from shallow subtidal zone with high
and low energy lagoonal environments. Intraclasts of mud-
stones without allochems and mudstones with fine parallel
laminations (<1 mm) originated mainly in the shallow subtidal
to supratidal area. Co-occurrence of these components in the
layers indicate redeposition of storm-eroded and transported
material. Poor sorting of the allochems with great differences
in their size, mixing of redeposited components of varied ori-
gin in fine-grained matrix is common.
Carbonate formation in both sections also provided field ev-
idence of lenslike shape, as well as sudden lateral changes of
microfacies in the beds. All of these observations allow us to
interpret the described carbonates as the sediments of channels
or depressions in the rugged relief of the peritidal area.
The claystones of the superposed formation above the Ram-
sau Dolomite Formation are well-sorted sediments with rare
internal textures. The boundary claystones on the base of the
Tržinovo Formation reveals the very fine parallel lamination,
where the laminae are sometimes enriched by epigenetic py-
rite grains and rare pyrite concretions, which indicate varia-
tions in the redox potential during the sedimentation. The
content of carbonate changes in the beds. Presence of the con-
chostracans and absence of a typical marine biota indicate
brackish up to freshwater conditions. The claystone sediments
could be deposited from mud suspensions, which overflow the
barrier from the channelized area filled by sandstone-rich sedi-
ments of the Lunz Beds. Like the Carnian terrestrial systems
of Europe, these channels were flooded in monsoonal epi-
sodes, then abandoned as pools with standing water and colo-
nized by Euestheria (cf. Simms & Ruffell 1990). Therefore, in
less humid phases the terrigenous claystones could alternate
again with peritidal carbonates.
The sedimentary area of the carbonate claystone deposition
in the Tržinovo Formation was separated from the terrigenous
clastic systems. The shales were deposited in shallow depres-
sions behind barriers which inhibited input of coarser materi-
al. The lens-shaped beds of the Tržinovo Formation represent
channels and depressions, which subsided due to tectonic dif-
ferentiation of the carbonate platform. Initial subsidence is al-
ready recorded in the underlying carbonates, which differ
from the surrounding dolomites by accumulation of lithoclasts
and bioclasts from different shallow-water environments. The
Lunz Beds close to A section do not exhibit cyclic organiza-
tion. It is supposed, that the unique development of the Car-
nian sediments in the Tržinovo Valley, reflects the specific
local conditions.
Dolomites of the Ramsau Formation cropped out in places
from below the bed No. 04 in road-cut section A. They are
bright grey in colour and locally contain beds of dolomitic
breccias to conglomerates (see Fig. 4c). Some of the litho-
clasts in these dolorudites exhibit the Liesegang bands (rusty-
red on weathered surfaces), which were formed before their
deposition in breccia beds (see Fig. 4d). Their origin is ex-
plained by redox gradients, when oxidizing and reducing flu-
ids met in an iron-enriched zone, that formed soon after
sediment deposition (e.g. Breit 2001). Genesis of the litho-
clasts with Liesegang bands is connected with phreatic envi-
ronments. The carbonate rudites have predominantly micritic
matrix, only with local cements. The individual lithoclasts
have subangular to well rounded shapes, frequently deformed
by stylolites, and between 0.07 to 20 mm in size. Breccias ex-
hibit squeezed and packed clast fabrics. We suppose that the
individual lithoclasts were derived from intertidal and su-
pratidal environments, whereas the clasts with Liesegang
bands come from the freshwater environments. Because of
imperfect size-sorting, mixing of lithoclasts of various origin
and erosional contact with the underlying bed the described
rudites seem to represent the storm lags (see Demicco & Hardie
1994 for comparison). In the second explanation, the breccias
could represent the basal high-energy infill of channel inci-
sion, formed in an erosive drainage system at the top of a sub-
aerially exposed platform during drops of relative sea level
(see Grélaud et al. 2010 for comparison).
Tržinovo Formation – new lithostratigraphic unit
On the basis of field works, lithological, microfacial, sedi-
mentological and paleontological research we suggest defin-
ing the Tržinovo Formation as a new lithostratigraphic unit.
Its name is derived from the Tržinovo Valley on the right
(western) site of the valley between Zázrivá and Párnica vil-
lages at the eastern tip of the Malá Fatra Mts. The new for-
mation oucrops in two localities. The stratotype and locus
typicus of the Tržinovo Formation occur in the forrest road
cut section with GPS coordinates N 49°13
’16” and
E 19°09
’24” (A-section see Fig. 1).
The lithological composition of the new formation marked-
ly differs from the presumably isochronous clastic sediments
of the Lunz Beds and other sediments of the Carnian forma-
tions of the Krížna Unit. The new formation consists of two
members: the Tržinovo Claystone with six layers of shales
(No. I—VI), and the Tržinovo Dolomite with interbeds of dolo-
mites and dolomitic limestones (see Fig. 3).
The Tržinovo Claystone is represented by grey, dark grey,
grey-yellowish, grey-greenish dolomitic claystones with shaly
desintegration and variable content of terrigenous clay frac-
tion (65.6 up to almost 100 %). In this fraction, illite strongly
predominates over kaolinite. The lowermost layer of claystone
(No. I) contains rare macrofossils including the brachiopods
Lingularia ex gr. tenuissima (Bronn) and conchostracans
129
SILICICLASTICS IN TRIASSIC DOLOMITE: CONSTRAINTS FOR CARNIAN PLUVIAL EVENT (W CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
Euestheria minuta (von Zieten), presented by Kozur &
Weems (2010). Both fossil groups were found in the lower
part of bed No. I. Brachiopods are more common than con-
chostracans, but their stratigraphical importance is low. Silt
quartz admixture of epigenetic pyrite grains was found very
scarcely in this basal finely laminated claystone.
The Tržinovo Dolomite Member is represented by pale grey
bedded dolomites and sporadic dolomitic limestones. These
carbonates are micritic, with rare allochems including peloids,
intraclasts (pelsparites, intramicrites) and finely-laminated
bindstones. Some beds have fenestral fabrics.
The lower boundary of the Tržinovo Formation is marked
by the lower bedding plane of bed No. I. This plane overlies
the dolomite bed No. 8a, belonging to the topmost bed of the
Ramsau Dolomite Formation, and that without any marks of
erosion. The upper boundary of the Tržinovo Formation cor-
responds to the upper bedding plane of the claystone layer
No. VI. This boundary claystone divided the Tržinovo For-
mation from the Hauptdolomit Formation. Thickness of the
Tržinovo Formation is approximately 12.3 m (see Fig. 3). The
parastratotype of this formation is only known in the B-sec-
tion at N 49°12
’36” and E 19°08’05” (Fig. 1). The new for-
mation has a lens-shaped prolongation, and is known only
from two above mentioned localities in the Malá Fatra Mts, up
to now.
A very similar formation of Lower Carnian sediments in the
Krížna Unit was detected in the Ružbachy Horst in Eastern
Slovakia by Kullmanová & Nemčok (1985). They described
small outcrop of a formation of dolomites and dolomitic clay-
stones about 1.5 m thick. The dolomites are similar in the
composition of the Carnian foraminifers to the association
from the Malá Fatra Mts. The dolomitic claystones near
Ružbachy also contain marine bivalves Costatoria/Costato-
ria/cf. goldfussi (Alberti in Zieten) and brachiopods of the ge-
nus Lingula in the claystones (Kochanová in Kullmanová
1974). On the basis of the bivalves Kullmanová & Nemčok
l.c. assigned this formation to the Cordevolian, whereas Polák
in Janočko et al. (2000) preferred its stratigraphic attribution
to the Julian and Tuvalian. Recently, the Ružbachy sections
are not uncovered for study.
Stratigraphic attribution of the Tržinovo Formation also
depends on the definition of the Carnian Stage. Recent de-
bate makes problematic the lower boundary of the Carnian
Stage, when the traditional base of the Cordevolian ( = base
of the A on Zone sensu Krystyn 1978) have been questioned
due to asynchronous FOs of Trachyceras species (Mietto &
Manfrin 1999). Therefore, Broglio Loriga et al. (1998) pro-
posed to shift the base of the Cordevolian to the base of
Daxatina canadensis Zone, which was the cosmopolitan am-
monoid species. Nevertheless, some Late Ladinian species
like Frankites are still present in the Cordevolian, which led
to assignment of the Daxatina canadensis Zone to the Ladin-
ian (see Mietto et al. 2007 and Kozur & Bachman 2010 for
summary). Consequently, the Carnian stratigraphy has been
improved by a two-stage subdivision into the Julian and Tu-
valian (e.g. Lucas 2010, etc.).
The age of the Tržinovo Formation can be approximated
on the basis of conchostracan species Euestheria minuta
(von Zieten), which is known from the Longobardian, but is
also still present in the entire Cordevolian = Julian 1/I (Kozur
& Bachmann 2010). This age is also supported by the Carnian
foraminifers from topmost carbonate beds of the Ramsau Dolo-
mite Formation, which is directly overlain by the Tržinovo
Formation (see below). Age control in deposion of the
Tržinovo Formation is related to the Carnian Pluvial Event,
which took place in the latest Julian 1/IIc or at the Julian/Tu-
valian boundary (Hornung et al. 2007a,b; Simms & Ruffell
1989; Rostási et al. 2010, etc.).
Paleontology and biostratigraphy
Upper Triassic sediments contain lingulide brachiopods and
conchostracans in the Tržinovo Formation and foraminifers in
the Ramsau Dolomite Formation. These fossils were studied
from the viewpoints of systematic description, ecology and
taphonomy and biostratigraphic importance.
Brachiopods
Order: Lingulida Waagen, 1885
Superfamily: Linguloidea Menke, 1828
Family: Lingulidae Menke, 1828
Lingularia Biernat & Emig, 1993
Lingularia ex gr. tenuissima (Bronn, 1837)
(Fig. 7a—e )
1830 Lingula Bronn – p. 128
1830 Lingula tenuissima n.s. Bronn – p. 230 (nomen nudum)
1837 Lingula tenuissima Bronn – p. 158, pl. 13, fig. 6b
1920 Lingula tenuissima Bronn – Diener, p. 16 (cum syn.)
1927 Lingula tenuissima Bronn – Ogilvie Gordon, p. 31, pl. 3, fig. 40
1934 Lingula tenuissima Bronn – Kirchner, p. 90, pl. 2, fig. 1
1935 Lingula tenuissima Bronn – Leonardi, p. 31, pl. 1, fig. 3—4
1958 Lingula tenuisima Bronn (sic) – Virgili, p. 534, fig. 62/4
?1960 Lingula cf. tenuissima Bronn – Šuf, pl. 14, fig. 7
1968 Lingula tenuissima Bronn – Nagy, pl. 2, fig. 3
1968 Lingula tenuissima Bronn – Broglio Loriga, p. 201, pl. 1, fig. 1—6
1972 Lingula tenuissima Bronn – Encheva, p. 19, pl. 9, fig. 1—6, pl. 10,
fig. 7—8
1985 Lingula tenuissima Bronn – Senkowiczowa, p. 24, p. 24, pl. 1,
fig. 2—3
1988 Lingula tenuissima Bronn – Siblík, p. 9 (cum syn.).
1991 Lingula tenuissima Bronn – Biely & Rakús, p. 6, pl. 1, fig. 4,7
1997 Lingula tenuissima Bronn, 1851 – Calzada & Magrans, p. 45,
fig. 1
1999 Lingula tenuissima Bronn – Sulser, p. 45, text-fig. 1
(synonymy includes the references with figured specimens only)
M a t e r i a l: 300 more or less fragmentary separated valves,
bed I, section A, Tržinovo Valley.
D e s c r i p t i o n: Elongate oval, slightly convex or nearly
flat separate valves with up to 18.5 mm in length and 9.5 mm
in width. Lateral margins slightly convex, anterior margin
rounded. Maximum width at the mid-length. Posterior margin
rounded but majority of valves with damaged umbonal parts.
Ornamentation consists of growth lines – concentric fila of
variable prominence and spacing. Due to damaged posterior
parts of most specimens, the distinguishing of pedicle and bra-
130
SÝKORA, SIBLÍK and SOTÁK
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
chial valves of our specimens was practically unfeasible. In-
ternal characters are not well visible. The interior of one pedi-
cle valve only showed narrowly V-shaped grooves joining
near the posterior adductor.
P r e s e r v a t i o n: Most specimens are preserved as more or
less fragmentary and depressed internal moulds of separate
valves and have thin black organophosphatic shell preserved
only in fragments. They are lying scattered, parallel to the
bedding. The character of preservation has not enabled infor-
mation on character states of interior soft parts. No size-sort-
ing of valves was ascertained.
R e m a r k s: Findings of Triassic lingulid brachiopods are
relatively rare and their preservation not very favourable.
However, the specimens may occur in great numbers (“Lingu-
la — Beds”). Mass occurrences of lingulids are reported from
several localities in the Italian Lower Triassic (“Scythian”) by
Fig. 7. Lingulid brachiopods and conchostracans from the Tržinovo Formation. a—e – Lingularia ex gr. tenuissima (Bronn); f – Euesthe-
ria minuta (von Alberti) from bed I, section A. Tržinovo Formation. Fossils are deposited in the Slovak National Museum Bratislava under
numbers Z 24741—Z 24745 (a—e) and Z 24746 (f).
Broglio Loriga et al. (1980). Those brachiopods were deter-
mined as Lingula tenuissima in former publications (e.g. Bro-
glio Loriga 1968) or Lingula sp. in the subsequent ones. These
authors (1980) also summarized lingulid findings in the Low-
er Triassic from all over the world. On the other hand, mass
occurrences in the Iberian Range in Spain were ascertained in
the Uppermost Muschelkalk (Cordevolian) by Márquez-Alia-
ga et al. (1999), and more recently by Márquez-Aliaga et al.
(2007). The identification of their respective specific appurte-
nances is difficult. As there is a very limited number of avail-
able external features, and the general outlines and external
features of the shell are variable and not sufficient for the spe-
cific identification, the determinations remain in many cases
doubtful. The shape was certainly influenced by environmen-
tal conditions and later by fossilization, which could change
shell convexity (thickness of valves).
131
SILICICLASTICS IN TRIASSIC DOLOMITE: CONSTRAINTS FOR CARNIAN PLUVIAL EVENT (W CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
Most of the past Triassic lingulid finds were assigned to
Lingula tenuissima Bronn, which was described from the Ger-
manic Triassic. However, due to frequent bad preservation,
some identifications remain questionable. Very similar lin-
gulids, also described from the Germanic Triassic Lingula ke-
uperea Zenker, 1834, Lingula calcaria Zenker, 1834 and
Lingula zenkeri v. Alberti, 1864 were synonymized by some
authors. The names of the first two of them could have priority
in the case of identity with L. tenuissima. However, Lingula
keuperea as figured by Zenker, 1834 shows in comparison to
L. tenuissima relatively larger, subtrigonal outline with maxi-
mum width in the anterior third of the shell. Lingula calcaria
seems to be a juvenile of L. keuperea. The Alpine species
Lingula christomani Skuphos, 1893 from the “Partnach-
schichten” and Lingula fischeri Suess, 1854 from the Hallstatt
Limestone were extremely rare when established, and have
not been ascertained since their description. They both would
deserve a revision.
In 1977 Pajaud assigned “tenuissima” to Glottidia Dall,
1870 on the basis of the presence of divergent laminae in pedi-
cle valve. This was adopted later also by Siblík (1983). How-
ever, Glottidia occurs in ?Cretaceous, Tertiary—Holocene
(Holmer & Popov 2000). This genus was quite recently con-
firmed in the Eocene in Antarctica (by Emig & Bittner 2005).
The usual generic assignment of the Triassic lingulids to
Lingula Brugui
è
re, 1797 was called in question recently by
Biernat & Emig (1993). They documented that Mesozoic lin-
gulid brachiopods differ significantly from their Recent rela-
tives, and established a new genus Lingularia, which occurs in
Carboniferous—Cretaceous and ?Tertiary formations. One of
their 3 new species, Lingularia siberica comes from the Mid-
dle Anisian of Siberia. According to the above mentioned au-
thors, the genera of the living lingulids probably arose in the
Early Cenozoic. The genus Lingula itself occurs within the
?Cretaceous, Tertiary and Holocene only (Holmer & Popov
2000, p. 36), its type species L. anatina Lamarck in the Ho-
locene. A detailed summary of the taxonomy and organization
of the Recent species of Lingula was given by Emig (1982,
2002). The external characters are important, but mainly fea-
tures of the shell interior are necessary for the differentiation
between Lingula and Lingularia, especially impression of
the pedicle nerve, shape of umbonal muscle scar etc. Howev-
er, the internal characters are ascertainable only rarely in the
Triassic fossil finds. According to Biernat & Emig (1993),
the shell of Lingularia externally differs from Lingula in its
less acute posterior margin and beak, and in its broadly
rounded anterior margin.
Most specimens from the Ve ká Lučivná Valley show a
more ovate outline in comparison to Bronn’s figure (1837),
which is very general, however. Their shell morphology, even
if fragmentary, is wholly consistent with the attribution to Lin-
gularia. Clearly comparable to them is the specimen of Lingu-
la tenuissima, figured by Calzada & Magrans (1997) from the
Upper Ladinian. The main synonymies of L. tenuissima are
presented here to show the large specific variability and distri-
bution. The possibility to verify the respective determinations
is in most cases unfeasible due to universal, short original de-
scriptions or insufficient illustrations. Also unclear muscle im-
prints can be easily misinterpreted in some cases. With regard
to the above mentioned uncertainties, the determination of our
material as Lingularia ex gr. tenuissima is considered reason-
able. It is quite possible, however, that so-called “tenuissima”
itself is not consistent and corresponds to several species.
D i s t r i b u t i o n: Lingularia tenuissima (Bronn) provides a
low biostratigraphic importance. This species was described
originally from the Buntsandstein and the Wellenkalk in the
Germanic Triassic (Bronn 1835—7). It occurs in the Lower to
Upper Triassic (Carnian) of the Germanic and Alpine do-
mains, and is reported from Germany, Spain, France (Vosges),
Switzerland, Netherland, Austria, Italy, Slovakia (SE Slova-
kia: e.g. Nandráž W of Jelšava, Rožňava, Muráň Plateau E of
Šumiac – all Early Triassic), Hungary, Bulgaria, Bosnia,
Greece, Israel, Algeria, Tunisia and China.
E c o l o g y a n d t a p h o n o m y: Lingulids are marine endo-
bionts, living mainly on offshore, shoreface and intertidal,
deltas and estuarine settings – soft substrates of finely
grained sand or clayey sand. They are tolerant of reduced
salinity – e.g. Robertson (1989) etc., of oxygen deficient set-
tings – e.g. Wignall & Hallam (1992), Schubert & Bottjer
(1995), and their short dwelling in fresh or brackish water is
possible. Kowalewski (1996) confirmed that recent lingulid
brachiopods have very low fossilization potential, and that
longer transport of their shells before fossilization is highly
improbable. After death they can undergo rapid mechanical
damage, and then disappear within weeks or months. Their
preservation in the fossil record may be enabled by a high rate
of sedimentation only or by catastrophy. On the basis of the
literature, Kowalewski l.c. concludes that the fossilization po-
tential in Paleozoic lingulids is higher than that in post-Paleo-
zoic ones. It seems that the claystones in section A have not
represented the normal substrate of lingulids (no presence of
the lingulid – derived trace fossils were found) but on the
contrary, they are responsible for their burial. An outline of
environmental, biological and taphonomic controls on lin-
gulid distribution was given in Zonnenveld et al. (2007).
Conchostracans
Conchostraca Sars, 1867
M a t e r i a l: A number of shells in bed I, section A,
Tržinovo Valley.
D e s c r i p t i o n: Conchostracans belong to the clam
shrimps, the systematic classification of which remains uncer-
tain (see Kozur & Weems 2010). Triassic conchostracans are
assigned to the suborder Spinicaudata Linder, 1945. Conchos-
tracan shells from the Tržinovo Formation reach sizes of
around 3 mm, they display expressive growth bands and chiti-
nous composition
(Fig. 7f).
S t r a t i g r a p h i c i m p o r t a n c e: The conchostracans from
bed No. I were originally determined as Euestheria cf. minuta
(von Zieten) by Kozur & Mock (1993). Their systematic attri-
bution to the species of Euestheria minuta (von Zieten) has
been improved by Kozur & Weems (2010). In the conchostra-
can zonation of Kozur & Weems (l.c.), this taxon is the nomi-
native species of the Euestheria minuta Zone, which has been
established in the Longobardian. The associations of this con-
è
132
SÝKORA, SIBLÍK and SOTÁK
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
chostracan zone are rather monospecific, different from those
found below in the Xiangxiella bicostata Zone and above in
the Laxitexella multireticulata Zone (Kozur & Weems, l.c.).
Nevertheless, the Late Ladinian conchostracan species of
Euestheria minuta was still presented in the entire Cordevo-
lian (Kozur & Bachmann 2010).
E c o l o g y a n d t a p h o n o m y: The occurrence and distri-
bution of the conchostracans are limited by conditions and cli-
mate (especially temperature, salinity and acidity). Webb
(1979), Kozur & Mock (1993) and Shen et al. (2002) men-
tioned that the ecological tolerance of the spinicaudatan crus-
taceans in Paleozoic was wider than that known from the
recent taxa. According to Kozur & Mock (l.c.), the conchost-
racans are restricted to freshwater to brackish environments
with salt concentration of about 10 ‰. They survived in plio-
haline environment with limiting salinity of 16.5 ‰. In recent
environments of Australia, the conchostracans do not with-
stand salinity above 5 ‰ (De Decker 1988). The conchostra-
cans proliferated in environments such as ponds mostly as
meiobenthic fauna. The chitinous shells of conchostracans are
very fragile and very sensitive to transportation. Therefore, the
shells are mostly found in sediments (mainly claystones) at
the places of their primary deposition in stagnant environ-
ments. Conchostracans are one-season animals. Their growth-
rate and frequency depends on temperature and nutrient
availability (see Webb 1979; Frank 1988, etc.). In conchostra-
Fig. 8. Foraminiferal microfauna from
terminal beds of the Ramsau Dolomite
Formation. 1—4 – Pilamminella kuthani
Salaj, index species of the Carnian fora-
miniferal biozone. 1—3 – sample bed. 0,
section A; 4 – sample bed 2, section A,
thin section No. 27290. 5—6 – Pilam-
minella cf. gemerica Salaj; 5 – sample
bed 0. section A; 6 – sample bed 2,
section B. Scale bar 200 m.
133
SILICICLASTICS IN TRIASSIC DOLOMITE: CONSTRAINTS FOR CARNIAN PLUVIAL EVENT (W CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
can-bearing bed No. I (section A), the shells of Euestheria oc-
cur together with valves of Lingularia, in which the lingulids
dominated. However, the total number of both species is quite
low, reflecting relatively unfavourable life conditions during
deposition. Therefore we suppose that these fossils represent
oryctocoenosis generated by postmortem transport of conchost-
racans and lingulids in suspension, by low-energy flows over
short distances. This flow regime is recorded by fine lamination
of fossil-bearing claystones. Similar thanatocoenoses of lin-
gulids and conchostracans are known as in the so-called “Esthe-
rienschichten” ( = “Estherienschiefer”) in the Germanic Keuper,
or in “Obererer Muschelkalk” (see Heller 1952; Merki 1961 for
summary). They are often reported as Lingula tenuissima Bronn
and Euestheria minuta (Goldfuss). In the latest correlation of
the Germanic Triassic with the international scale by Kozur &
Bachmann (2005, Fig. 7), the Upper Grabfeld Formation
(“Estherienschichten”) is assigned to the Cordevolian.
Foraminifers
M a t e r i a l: Foraminiferal microfauna from the limestones
in the upper part of the Ramsau Dolomite Formation
(Beds 1, 1a, 2, 2a – section A; 2 – section B). See distribu-
tion of foraminifers in Table 1.
Fig. 9. Foraminiferal micro-
fauna from terminal beds of
the Ramsau Dolomite Forma-
tion. 1—4 – Triadodiscus
eomesozoicus (Oberhauser),
type species of the Carnian in-
volutinids. 1 – sample bed 2,
section B; 2 – sample bed 2,
section
A,
thin
section
No. 27290; 3 – sample base
bed 1b, section A. 5 – Aulo-
tortus sinuosus Weynschenk,
sample bed 2, section A, thin
section No. 27291. 6—8 – Au-
lotortus praegaschei (Koehn-
Zaninetti). 6, 7 – sample
bed 1, section A, thin section
No. 27287b; 8 – sample
bed 2, section A, thin section
No. 27291. Scale bar 200 m.
134
SÝKORA, SIBLÍK and SOTÁK
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
D e s c r i p t i o n: The limestones contain an abundant micro-
fauna of ammodiscid, involutinid and various elongated and
multilocular foraminifers. The most typical ammodiscid fora-
minifers belong to the species Pilamminella kuthani Salaj
(Fig. 8.1—4), which represents the index species of the Carnian
biozone (Salaj 1969; Salaj et al. 1983). Some club-shaped for-
aminifers are closer to another Carnian species Pilamminella
cf. gemerica Salaj (Fig. 8.5—6) and Rectopilammina (Recto-
glomospira) senecta Trifonova. Involutinid foraminifers are
more diversified, including the species of Triadodiscus eome-
sozoicus (Oberhauser), Aulotortus praegaschei (Koehn-Za-
ninetti), Prorakusia salaji Di Bari & Laghi, Aulotortus
broennimanni Salaj and Aulotortus sinuosus Weynschenk
(Fig. 9). Trochospiral involutinids and nodosariids, rich asso-
Fig. 10. Foraminiferal microfauna from terminal beds of the Ramsau Dolomite Formation. 1 – Lamelliconus turris (Frentzen), sample bed 1,
section A, thin section No. 27287b; 2 – Multiseptida elongata Salaj, sample bed 2, section A; 3 – Nodosaria ordinata Trifonova, sample
bed 1, section A, thin section No. 27287b; 4 – Rectoglomospira (Rectopilammina) senecta Trifonova, sample bed 1, section A, thin section
No. 27287b; 5 – Meandrospira sp., sample bed 1, section A, thin section No. 27287; 6 – Ophthalmidium exiguum Koehn-Zaninetti, sample bed 1,
section A; 7 – Endothyra gruenbachensis Salaj, sample bed 0, section A, thin section No. 27287; 8 – Oberhauserella mesotriassica (Oberhauser),
sample bed 2, section A; 9 – Duostomina cf. alta Kristan-Tollmann, sample bed 2, section A, thin section No. 2729. Scale bar 100 µm.
135
SILICICLASTICS IN TRIASSIC DOLOMITE: CONSTRAINTS FOR CARNIAN PLUVIAL EVENT (W CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
ciations of which occur in the Late Ladinian—Carnian lime-
stones, are represented by Lamelliconus turris (Frentzen),
Nodosaria ordinata Trifonova and Multiseptida elongata
Salaj, Borza & Samuel (Fig. 10). The so-called Triassic
“globigeriniids” are represented by the species of Ober-
hauserella mesotriassica (Oberhauser) and Duostomina cf.
alta Kristan-Tollmann. The foraminiferal association is com-
pleted by the species of Endothyra gruenbachensis Ober-
hauser, Abriolina cf. mediterranea Luperto, Meandrospira
sp., Ophthalmidium exiguum Koehn-Zaninetti, etc.
S t r a t i g r a p h i c i m p o r t a n c e: The association of the
listed species allows us to determine the Carnian age of the
limestones. This age is indicated by three important species –
Pilamminella kuthani Salaj, Triadodiscus eomesozoicus
(Oberhauser) and Lamelliconus turris (Frentzen). The associa-
tion of these species is indicative for the Carnian formations
(cf. Ciarapica & Zaninetti 1983, 1984a,b; Peybernes et al.
1991; Fréchengues & Peybernes 1991; Fréchengues et al.
1993; Di Bari & Laghi 1994; Kolar-Jurkovšek et al. 2005,
etc.). The Carnian age is also supported by the species Aulo-
tortus praegaschei (Koehn-Zaninetti), Abriolina cf. mediter-
ranea Luperto, Multiseptida arcata Salaj, Borza & Samuel,
Rectoglomospira (Rectopilammina) senecta Trifonova and
Endothyra gruenbachensis Salaj.
E c o l o g i c i m p l i c a t i o n s: Foraminiferal microfauna al-
lows us to propose certain characteristics of the paleoeco-
logical conditions. Involutinid genera, like Triadodiscus,
Aulotortus, Lamelliconus, etc., belong to the lamellar arago-
nitic foraminifers, which are characteristic of high-energy and
shallow-water conditions. On the contrary, the agglutinated
benthic foraminifers, like Pilamminella, Rectoglomospira,
Meandrospira, etc., indicate the restricted low-energy envi-
ronments. The low-energy facies of micritic limestones re-
veals an abundance of elongated morphotypes like Nodosaria,
Pseudonodosaria, Frondicularia, etc., which could indicate
dysoxic conditions (cf. Ciarapica et al. 1987; Maurer & Ret-
tori 2002). Under a stress life condition, some nodosariids lost
the calcareous hyaline walls, exhibiting a preference for ag-
glutination (Multiseptida). These limestones are also enriched
in some epiplanktonic forms like the zoospores Globochaete
and juvenile lamellibranchians (“filaments”).
Discussion and conclusions
The Upper Triassic development of the Germanic and North
Tethyan shelf basins was significantly influenced by overall
siliciclastic input, which suffocated a carbonate factory and
led to the so-called “Carnian Crisis” (Hornung et al. 2007a),
Reingraben or Raibl Turnover (Schlager & Schöllnberger
1974), Carnian Pluvial Event (Simms & Ruffell 1989), Mid-
dle Carnian wet intermezzo (Kozur & Bachmann 2010) and
other related events. These Carnian terrigenous inputs reflect-
ed climatic warming, increased humidity, monsoonal precipi-
tation, enhanced weathering, extreme seasonality and
development of large fluvial systems (Simms & Ruffell 1990;
Krystyn 1991; Vissher et al. 1994; Mutti & Wissert 1995;
Hornung 2007; Hornung et al. 2007a,b; Roghi et al. 2010;
Kozur & Bachmann 2010, etc.). During the Carnian, these
Tethys-wide changes resulted in the demise of carbonate plat-
forms, enhanced input of siliciclastics, freshwater influx and
brackishing of shallow-marine basins, anoxic conditions, de-
crease of faunal diversity, biotic extinctions, etc. (later cit.).
The Carnian Crisis is documented between the uppermost
Aonoides Zone and uppermost Austriacum Zone, which corre-
spond to its duration between Julian 1/IIc and Julian 2/II (Hor-
nung & Brandner 2005; Hornung et al. 2007b). Subsequent
deposition of the Lunz Beds (s.s.) took place during Julian
2/IIa—2IIb, estimated at about 1 Myr duration. As the same
time as in the Northern Calcareous Alps, the Carnian Crisis is
Table 1: Distribution of foraminifers.
Bed numbers — Section A
Sec. B
Foraminiferal species
03 0
1
27287a
1
27287b
1
1b
base bed
1b
middle part
of bed
2
2
27403
2a
27290
2b
27289F
2
Microfacial type
W/G
W/P/G
G G P/G P
P P/G BM P/G P/G P/G
Pilamminella kuthani
R
C R F F C C R – C C R
Pilamminella gemerica
–
C –
R R R
–
–
–
R R –
Rectopilammina senecta
–
R –
R –
–
–
–
–
–
–
–
Triadodiscus eomesozoicus
C
C
R
R
– R R C R R R F
Aulotortus praegaschei
– R C R C C
F
R –
C
C R
Aulotortus sinuosus
R
C F C F R
F C R R R R
Aulotortus broennimanni
– –
C – C R C –
–
R R R
Lamelliconus turris
R –
R –
–
–
C R –
–
–
R
Nodosaria ordinata
C C R C
C R R F F F R F
Multiseptida elongata
R C R F C –
–
F C C R C
Oberhauserella mesotriassica
R R – –
–
–
–
R –
–
–
–
Duostomina alta
R R –
–
–
–
–
R –
–
–
–
Endothyra gruenbachensis
R C –
–
–
–
–
– –
–
–
–
Meandrospira sp.
– R R R –
–
R R –
–
–
–
Ophthalmidium exiguum
C – R R R –
–
R R R C R
Glomospirella sp.
R
R C C C C C C R C R C
Aulotortus sp.
C
C F C C R C R R C C C
Semiinvolutina sp.
R R C –
–
–
R –
–
–
R –
Prorakusia sp.
R
R C – R R R R – C C R
Explanation: R — rare, C — common, F — frequent, W — wackestone, P — packstone, G — grainstone.
136
SÝKORA, SIBLÍK and SOTÁK
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
also recorded in the Western Carpathians. Nevertheless, the
decreased rate of terrigenous input in the Western Carpathians
implies their more distant position from clastic sources than
the Northern Calcareous Alps.
Sedimentary changes in the Upper Triassic carbonate plat-
form of the Western Carpathians are manifested mainly from
the Carnian (Mahe et al. 1968; Andrusov & Samuel (Eds.)
1985). Manifestations of the most conspicuous changes are
known from the Hronic and Silicic Zones (Choč Unit, Jablonica
Unit), where the Carnian siliciclastic inputs are recorded by
the Reingraben Shales and Lunz Beds. Their thick formation
was described, for example, from the Dobrá Voda DV-1 and
Kuklov 3 boreholes (Michalík et al. 1992; Masaryk & Lint-
nerová 1997), where the black shales with halobiids are equiv-
alent to the Reingraben ( = Halobia) Shales, and to the Lower
Carnian event in full-marine environment and sabkha-type en-
vironment (Opponitz Formation). An enhanced input of silici-
clastics is also recorded in the intraplatform basins of the
Hronic Zone, where the Reifling limestones were replaced by
the Aonian Beds (Svarín Beds) and higher by the thick se-
quence of the Lunz Beds (Marschalko & Pulec 1967;
Michalík 1988; Masaryk et al. 1993; Havrila 1993). These
changes, which correspond to the Julian, were dated by the oc-
currence of Trachyceras aonoides Mojsisovics, Monophyllites
simoyi Hauer, Carnites floridus (Wulfen) and others am-
monoids from the transition beds between the Reifling Lime-
stones and Lunz Beds (Andrusov et al. 1973; Bystrický in
Andrusov & Samuel 1985). Palynological study provided the
Carnian spore-pollen patterns of the Lunz Beds (Planderová
1972). The Lunz Beds also occur in the Fatric Zone, where
they are developed as a thinner formation with sandstone-
dominated facies. The thickness of the Lunz Beds with “Aon
Schiefer” in the NCA is about 500 m (Behrens 1972) and in
the Choč Unit about 300 m (Marschalko & Pulec 1967). Their
thickness in the Krížna Unit is reduced perhaps to 50—80 m,
and in the Malá Fatra Mts about 10 m. In the Tatricum, the
Carnian siliciclastic sediments are infrequent, known up to
now only from the Ve ká Fatra Mts, from where Planderová &
Polák (1976) described the dark grey shales and marly clay-
stones with abundant pollen/spore assemblages.
The Tržinovo Formation newly described from the Malá
Fatra Mts also represents the siliciclastic systems of the Fatric
basin, but those from shallow-lagoonal up to not fully marine
environments. The presence of pliohaline and euryhaline fau-
na like conchostracans and lingulids point to brackish water
environments of the Tržinovo Formation. The claystone sedi-
ments of this formation could be deposited from mud suspen-
sions overspilled from submarine channels filled by the
coarser terrigenous sediments of the Lunz Beds. The Flood
basins of the Tržinovo Formation were rather shallow intra-
platform depressions with restricted extent (1.5 km between
two border sections A—B). Similar flood basins with siliciclas-
tics within the shalow-marine platform environments were
formed during the Carnian Pluvial Event also in the Dolomites
(Breda et al. 2009). Like the Carnian terrestrial systems of Eu-
rope, these inundation basins were flooded in monsoonal epi-
sodes, and than abandoned as pools with standing water and
colonized by Euestheria (cf. Simms & Ruffell 1990). There-
fore, in less humid phases the terrigenous claystone deposition
was suppressed, which led to the recovery of full-marine
conditions and precipitation of carbonates. Alteration of such
conditions during the Carnian Pluvial Episode is recorded by
the mixed siliciclastic-carbonatic sequence of the Tržinovo
Formation, and related similar sediments of the Krížna Units
(e.g. dolomitic claystones with Costatoria cf. goldfussi in
the Ružbachy Horst, Eastern Slovakia – Kullmanová &
Nemčok 1985).
Acknowledgments: The research has been supported by
VEGA Agency (2/0140//09 and 1/0274/10) and funds re-
ceived througth the Centre of Excellence for Integrative Re-
search of the Earth’s Geosphere (ITMS 26220120064,
European Regional Development Fund). The study of brachi-
opods was done within the framework of the Project 205/03/
1123 of the Grant Agency of the Czech Republic and of the
Research Program AVOZ30130516. We extend our thanks to
Eva Šamajová (Bratislava) for X-Ray analyses, Roman
Aubrecht and Jozef Hók (UK Bratislava) for help in transla-
tion and in graphic arrangement. Many thanks for critical re-
views of the manuscript to Gloria Ciarapica (Perugia),
Christian C. Emig (Marseille) and Jozef Michalík (Bratislava).
References
Andrusov D. & Samuel O. (Ed.) 1985: Stratigraphical dictionary of
Western Carpathians 2, L/Z. GÚDŠ, Bratislava, 1—359 (in Slo-
vak).
Behrens M. 1972: Schwermineralverteilungen und Sedimentstruk-
turen in den Lunzer Schichten (Karn, Trias, Österreich). Jb.
Geol. B.—A. 116, 51—83.
Biely A., Bujnovský A., Vozárová A., Klinec A., Miko O., Vozár J.,
Beňuška P., Bezák V., Hanzel V., Kubeš P., Liščák P., Lukáčik
E., Maglay J., Molák B., Pulec M., Putiš M. & Slavkay M. 1997:
Explanations to the geological map of the Low Tatra Mts.
1 : 50,000. GÚDŠ, Bratislava, 1—232 (in Slovak).
Biernat G. & Emig C.C. 1993: Anatomical distinctions of the Meso-
zoic lingulide brachiopods. Acta Palaeont. Pol. 38, 1/2, 1—20.
Breda A., Preto N., Roghi G., Furin S., Meneguolo R., Ragazzi E.,
Fedele P. & Gianolla P. 2009: The Carnian Pluvial Event in the
Tofane area (Cortina D’Ampezzo, Dolomites, Italy). Geo. Alp 6,
80—115.
Breit G.N. 2001: Early diagenetic ferric oxide accumulations formed
along redox gradients: examples from modern and ancient flu-
vial sedimentary units. Abstracts, GSA Annual Meeting Boston,
33, 6, A281.
Broglio Loriga C. 1968: Some considerations about Lingula tenuis-
sima Bronn from the Werfenian Fm. of the Dolomites. Ann.
Univ. Ferrara (N.S.) 9, 4, 12, 189—202 (in Italian).
Broglio Loriga C., Neri C. & Posenato R. 1980: The Lingula zone of
the Scythian (Lower Triassic). Stratigrafia e paleoecologia.
Ann. Univ. Ferrara (N.S.) 9, 6, 6, 91—130 (in Italian).
Broglio Loriga C., Cirelli S., De Zanche V., di Bari D., Gianolla P.,
Laghi G.F., Lowie W., Manfrin S., Mastandrea A., Mietto P.,
Muttoni G., Neri C., Posenato R., Reichichi M., Rettori R. &
Roghi G. 1998: The Prati di Stuores/Stuores Wiesen section
(Dolomites, Italy): a candidate Global Stratotype Section and
Point for the base of the Carnian stage. Riv. Ital. Paleont. Stratigr.
105, 1, 37—78.
Bronn H.G. 1830: Gaea Heidelbergensis. 1—237, Heidelberg, Leipzig.
Bronn H.G. 1835—7: Lethaea geognostica I. (Atlas 1837, Pl. 1—47,
Stuttgart), 1—544.
137
SILICICLASTICS IN TRIASSIC DOLOMITE: CONSTRAINTS FOR CARNIAN PLUVIAL EVENT (W CARPATHIANS)
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
Calzada S. & Magrans J. 1997: One new finding of Lingula from the
Triassic of the Pyrenees. Batalleria 7, 45—46 (in Italian).
Ciarapica G. & Zaninetti L. 1983: Faune a Foraminiferes Ladino-
Carniens dans les Schistes de Fornovolasco, “Unita delle Panie”
(Alpes Apuanes, Italie). Rev. Paléobiologie 2, 1, 47—59.
Ciarapica G. & Zaninetti L. 1984a: Foraminiferes et biostratigraphie
dans le Trias supérieur de la série de la Spezia (Dolomies de
Coregna et Formation de la Spezia, nouvelles formations), Ap-
ennin septentrional. Rev. Paléobiologie 3, 1, 117—134.
Ciarapica G. & Zaninetti 1984b: Aulotortus praegashei (Koehn-Za-
ninetti 1968): revision taxonomique et stratigraphique sur la
base du materiel-type. Rev. Paléobiologie 3, 1, 53—61.
Ciarapica G., Cirilli S., D’Argenio B., Marsella E., Passeri L. & Za-
ninetti L. 1987: Late Triassic open and euxinic basins in Italy.
Rend. Soc. Geol. Ital. 9, 157—166.
De Decker P. 1988: Biological and sedimentary facies of Australian salt
lakes. Palaeogeogr. Palaeoclimatol. Palaeoecol. 62, 237—270.
Demicco R.V. & Hardie L.A. 1994: Sedimentary structures and early
diagenetic features of shallow marine carbonate deposits. Soc.
Econ. Paleontol. Mineral., Atlas Ser. 1, 1—255.
Di Bari D. & Laghi G.F. 1994: Involutinidae Bütschli (Foraminifer-
ida) in the Carnian of the Northeastern Dolomites (Italy). Mem.
Sci. Geol. 46, 93—118.
Emig C.C. 1982: Taxonomie du genre Lingula (Brachiopodes, Inar-
ticulés). Bull. Mus. Hist. Nat. Paris (4) 4 A, 3/4, 337—367.
Emig C.C. 2002: Death: a key information in marine palaeoecology.
Col. Lecio Encontres 5, 21—26.
Emig C.C. & Bitner M.A. 2005: Glottidia (Brachiopoda: Lingulidae)
from the Eocene La Meseta Formation, Seymour Island, Antarc-
tica. Palaeontology 48, 423—431.
Flügel E. 2004: Microfacies of carbonate rocks, analysis, interpreta-
tion and application. Springer, Berlin, Heidelberg, New York,
1—976.
Frank P.W. 1988: Conchostraca. Palaeogeogr. Palaeoclimatol. Palae-
oecol. 62, 399—403.
Fréchengues M. & Peybernes B. 1991: Associations de foraminiferes
benthiques dans le Trias carbonaté (Anisien, Ladinien-Carnien et
Rhétien) des Pyrénées Espagnoles. Acta Geol. Hisp. 26, 1, 67—73.
Fréchengues M., Peybernes B., Martini R. & Zaninetti L. 1993: Les
foraminiferes benthiques Ladino-Carniens du “Muschelkalk”
des Pyrénées Francaises a L’Est de la Garonne. Rev. Micropal.
36, 2, 111—120.
Grélaud C., Razin P. & Homewood P. 2010: Channelized systems in
an inner carbonate platform setting: differentiation between inci-
sions and tidal channels (Natih Formation, Late Cretaceous,
Oman). In: van Buchem F.S.P., Gerdes K.D. & Esteban M.
(Eds.): Mesozoic and Cenozoic Carbonate Systems of the Medi-
terranean and the Middle East: Stratigraphic and diagenetic ref-
erence models. Geol. Soc. London, Spec. Publ. 329, 163—186.
Haško J. & Polák M. 1979: Explanations to the geological map of
the Kysucké Vrchy Hills and Kriváň part of the Malá Fatra
Mts. 1 : 50,000. GÚDŠ, Bratislava, 1—145 (in Slovak, English
summary).
Havrila M. 1993: The research of the basinal and slope sediments of
the Biely Váh succession and paleogeography of the Hronicum.
Manuscript, GÚDŠ, Bratislava, 1—49 (in Slovak).
Heller F. 1952: Die stratigraphische Verbreitung und die Bedeutung
des Vorkommens von Lingula und Estheria im fränkischen Keu-
per. Geol. Blätt. NO-Bayern 2, 57—63.
Holmer L.E. & Popov L.E. 2000: Lingulida. In: Kaesler R.L. (Ed.):
Treatise on invertebrate paleontology, pt. H – Brachiopoda, re-
vised. Boulder, Colorado and Lawrence, Kansas 2, 32—97.
Hornung T. 2007: Multistratigraphy of the Draxllehen Quarry near
Berchtes-Gaden (Tuvalian—Lacian 2). Implications for Hallstatt
Limestone sedimentation and palaeoclimate in the aftermath of
the “Carnian crisis”. Austrian J. Earth. Sci. 100, 82—99.
Hornung T. & Brandner R. 2005: Biochronostratigraphy of the Rein-
graben Turnover (Hallstatt Facies Belt): Local black shale
events controlled by regional tectonics, climatic change and
plate tectonics. Facies 51, 460—479.
Hornung T., Brandner R., Krystyn L., Joachimski M.M. & Keim L.
2007a: Multistratigraphic constraints on the NW Tethyan “Car-
nian Crisis”. In: Lucas S.G. & Spielmann J.A. (Eds.): The Glo-
bal Triassic. New Mexico Mus. Nat. Hist. Sci. Bull. 41, 59—67.
Hornung T., Krystyn L. & Brandner R. 2007b: A Tethys-wide mid-
Carnian (Upper Triassic) carbonate productivity crisis: Evidence
for the Alpine Reingraben Event from Spiti (Indian Himalaya)?
J. Asian Earth Sci. 30, 2, 285—302.
Janočko J., Gross P., Polák M., Potfaj M., Jacko S. (Jun.), Rakús M.,
Halouzka R., Jetel J., Petro ., Kubeš P., Buček S., Köhler E.,
Siráňová Z., Zlinská A., Halásová E., Hamršmíd B., Karoli S.,
Žec B., Fejdiová O., Milička J., Boorová D. & Žecová K. 2000:
Explanations to geological map of the Spišská Magura Region.
ŠGÚDŠ, Bratislava, 1—174 (in Slovak, English summary).
Kobluk D.R. & Risk M.J. 1977: Calcification of exposed filaments of
endolithic algae, micrite envelope formation and sediment pro-
duction. J. Sed. Petrology 47, 2, 517—528.
Kolar-Jurkovšek T., Gazdzicki A. & Jurkovšek B. 2005: Conodonts
and foraminifera from the “Raibl-type” (Carnian) of the Kara-
vanke Mountains, Slovenia: stratigraphical and paleobiological
implications. Geol. Quart. 49, 4, 429—438.
Kowalewski M. 1996: Taphonomy of a living fossil: The lingulide
brachiopod Glottidia palmeri Dall from Baja California, Mexi-
co. Palaios 11, 3, 244—265.
Kozur H.W. & Bachmann G.H. 2005: Correlation of the Germanic
Triassic with the international scale. Albertiana 32, 21—35.
Kozur H. & Bachmann G.H. 2010: The Middle Carnian wet inter-
mezzo of the Stuttgart Formation (Schilfsandstein), Germanic
Basin. Palaeogeogr. Palaeoclimatol. Palaeoecol. 290, 107—119.
Kozur H.W. & Mock R. 1993: The importance of conchostracans for
the correlation of continental and marine beds. In: Lucas S.G. &
Morales M. (Eds.): The nonmarine Triassic. New Mexico Mus.
Nat. Hist. Sci. Bull. 3, 261—266.
Kozur H.W. & Weems R.E. 2010: The biostratigraphic importance of
conchostracans in the continental Triassic of the northern hemi-
sphere. Geol. Soc. London, Spec. Publ. 334, 315—417.
Krystyn L. 1978: Eine neue Zonengliederung im alpin mediterranen Un-
terkarn. In: Zapfe H. (Ed.): Beiträge zur Biostratigrafie der Tethys-
Trias. Schr. Erdwiss. Komm. Österr. Akad. Wiss. 4, 37—75.
Krystyn L. 1991: Die Fossillagerstätten der Alpinen Trias. Excur-
sionführer, Universität Sien, 1—61.
Kullmanová A. 1974: Lithological and microfacial research of
Ružbachy Mesozoikum, Manuskript. Geofond, No. 33669, Bra-
tislava, 1—191 (in Slovak).
Kullmanová A. & Nemčok J. 1985: Geology of Ružbachy Spa. Geol.
Práce, Spr. 82, 89—110 (in Slovak, English summary).
Leonardi P. 1935: Lower Triassic in the area of Tre Venezie. Mem.
Istit. Geol. R. Univ. Padova 11, 1—136, 3 figs., 8 pls. (in Italian).
Lexa J., Bezák V., Elečko M., Eliáš M., Konečný V., Less Gy.,
Mandl G.W., Mello J., Pálenský P., Pelikán P., Polák M., Potfaj
M., Radócz Gy., Ryłko W., Schnabel G.W., Stráník Z., Vass D.,
Vozár J., Zelenka T., Biely A., Császár G., Čtyroký P., Kaličiak
M., Kohút M., Kovács S., Mackiv B., Maglay J., †Nemčok J.,
Nowotny A., Pentelényi L., Rakús M. & Vozárová A. 2000:
Geological map of Western Carpathians and adjacent areas
1 : 500,000. Ministry of the Environment of Slovak Republic,
Geol. Surv. of Slovak Rep., Štátny geologický ústav D. Štúra
with coop., Geol. Surv. of Austria, Czech Geol. Surv., Geol. Inst.
of Hungary, Polish Geol. Inst., Geol. Surv. of Ukraine.
Lucas S.G. 2010: The Triassic timescale: an introduction. In: Lucas
S.G. (Ed.): The Triassic timescale. Geol. Soc. London, Spec.
Publ. 334, 1—16.
138
SÝKORA, SIBLÍK and SOTÁK
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA
GEOLOGICA CARPATHICA, 2011, 62, 2, 121—138
Mahe M. 1986: Geological Structure of the Czechoslovak Car-
pathians. Part 1. Paleoalpine Units. VEDA, SAV, Bratislava, 1—503
(in Slovak).
Mahe M., Buday T., Hanzlíková E., Chmelík F., Kamenický J.,
Koráb T., Kuthan M., Matějka A., Nemčok J., Pícha F., Roth
Z., Seneš J., Scheibner E., Stráník Z., Vaškovský I. & Žebera
K. 1968: Regional Geology of Czechoslovakia. Part II. The
West Carpathians. Geol. Surv., Czechoslovakia, Academia,
Praha, 1—723.
Mahe M., Kamenický J., Fusán O. & Matějka A. 1967: Regional ge-
ology of Czechoslovakia. Part 2, 1. The West Carpathians.
ÚÚG, ČSAV, Praha, 1—496 (in Slovak).
Márquez-Aliaga A., Emig C.C. & Brito J.M. 1999: Triassic lingulide
brachiopods from the Iberian Range (Spain). Géobios 32, 6,
815—821.
Márquez-Aliaga A., Emig C.C. & López-Gómez J. 2007: Triassic
Lingularia (Brachiopoda) from Moya (SE Iberian Ranges,
Spain). XXIII Jornadas de Paleontologia, Caravaca de la Cruz,
Murcia, Résum, 121—122.
Marschalko R. & Pulec M. 1967: Sedimentology of the Lunz beds.
Geol. Zborn. 18, 2, 331—344.
Masaryk P. & Lintnerová O. 1997: Diagenesis and porosity of the
Upper Triassic carbonates of the pre-Neogene Vienna Basin
basement. Geol. Carpathica 48, 6, 371—386.
Masaryk P., Lintnerová O. & Michalík J. 1993: Sedimentology,
lithofacies and diagenesis of the sediments of the Reifling in-
traplatform basins in the Central Western Carpathians. Geol.
Carpathica 44, 4, 233—249.
Maurer F. & Rettori R. 2002: Middle Triassic foraminifera from the
Secada core (Dolomites, Northern Italy). Riv. Ital. Paleont.
Stratigr. 108, 3, 391—398.
Merki P. 1961: Der obere Muschelkalk im östlichen Schweizer Jura.
Eclogae Geol. Helv. 54, 1, 137—219.
Michalík J. 1988: Svarín locality. In: Excursion guide of the Paleon-
tologica conference in Mi ava near Nižné Ružbachy. D. Štúr
Geol. Inst., Bratislava, 17—21.
Michalík J., Broska I., Franců, Jendrejáková O., Kochanová M.,
Lintnerová O., Masaryk P., Papšová J., Planderová E., Šucha
V. & Zatkalíková V. 1992: The borehole Dobrá Voda DV-1
(1140.8 m, Konča Skaliek) in Brezovské Karpaty Mts. Region.
Geol. Západ. Karpát 27, 7—139.
Mietto P. & Manfrin S. 1999: A debate on the Ladinian-Carnian
boundary. Albertiana 22, 23—27.
Mietto P., Andreetta R., Broglio Loriga C., Buratti N., Cirilli S., De
Zanche V., Furin S., Gianolla P., Manfrin S., Muttoni G., Neri
C., Nicora A., Posenato R., Preto N., Rigo M., Roghi G. & Spötl
C. 2007: A candidate of the global stratotype section and point
for the base of the Carnian Stage (FAD of Dexatina) in the Prati
di Stuores/Stuores Wiesen section (southern Alps, NE Italy).
Albertiana 36, 78—97.
Mutti M. & Weissert H. 1995: Triassic monsoonal climate and its
signature in Ladinian—Carnian carbonate platforms (Southern
Alps, Italy). J. Sed. Res. B65, 3, 357—367.
Pajaud D. 1977: Choix et hiérarchisation des caract
è
res taxinomiques
pour la reconaissance des Lingules post-paléozoiques. Géobios
10, 6, 961—965.
Peybernes B., Martini R. & Zaninetti 1991: Les Foraminiferes
benthiques du Trias carbonaté (Ladinien—?Carnien et Rhétien de
Corse). Geobios 24, 6, 683—696.
Planderová E. 1972: A contribution to palynological research of
Lunz Beds in West-Carpathians Region. Geol. Práce, Spr. 58,
55—77.
Planderová E. & Polák M. 1976: On age of Triassic dolomitic and
shaly beds in Tatric Unit of Ve ká Fatra Mts. Geol. Práce, Spr.
65, 75—79 (in Slovak, English summary).
Rakús M., Elečko M., Gašparík J., Gorek J., Halouzka R., Havrila
M., Horniš J., Kohút M., Kysela J., Miko O., Pristaš J., Pulec
M., Vozár J., Vozárová A. & Wunder D. 1988: Geological map
of the Lúčanská Malá Fatra Mts. 1 : 50,000. SGÚ, GÚDŠ, Bra-
tislava.
Robertson J.D. 1989: Physiological constraints upon marine organ-
isms: Trans. Roy. Soc. Edinburgh, Earth Sci. 80, 225—234.
Roghi G., Gianolla P., Minarelli L., Pilati C. & Preto N. 2010: Pa-
lynological correlation of Carnian humid pulses throughout
western Tethys. Palaeogeogr. Palaeoclimatol. Palaeoecol. 290,
1—4, 89—106.
Rostási A., Raucsik B. & Varga A. 2010: Paleoenvironmental controls
on the clay mineralogy of Carnian sections from the Transdanubi-
an Range (Hungary). Palaeogeogr. Palaeoclimatol. Palaeoecol.
doi: 10.1016/j.palaeo.2010.12.013
Salaj J. 1969: Essai de Zonation dans le Trias des Carpates Occiden-
tales d’apres les Foraminiferes. Geol. Práce, Spr. 48, 123—128.
Salaj J., Borza K. & Samuel O. 1983: Triassic foraminifers of the
West Carpathians. Geol. Ústav D. Štúra, 9—215, Plat. I-CLVII.
Schlager W. & Schöllnberger W. 1974: Das Prinzip stratigraphischer
Wenden in der Schichtfolge der Nördlichen Kalkalpen. Mitt.
Geol. Gesell. Wien 66/67, 165—193.
Schubert J.K. & Bottjer D.J. 1995: Altermath of the Permian—Trias-
sic mass extinction event: Paleoecology of Lower Triassic car-
bonates in the western USA. Palaeogeogr. Palaeoclimatol.
Palaeoecol. 116, 1—39.
Shen Y., Garassino A. & Teruzzi G. 2002: Studies on Permo-Trias of
Madagascar 4. Early Triassic conchostracans from Madagascar.
Atti Soc. Ital. Sci. Natur. Mus. Civ. Storia Nat. 143, 1, 3—11.
Siblík M. 1983: Triassic brachiopods of the Northern Alps and their
generic attributions. Schriftenr. Erdwiss. Komm. Österr. Akad.
Wiss. 5, 277—286.
Simms M.J. & Ruffell A.H. 1989: Synchroneity of climatic change
and extinctions in the Late Triassic. Geology 17, 265—268.
Simms M.J. & Ruffell A.H. 1990: Climatic and biotic change in the
Late Triassic. J. Geol. Soc. London 147, 321—327.
Sýkora M. 2003: Skeletal and detrital limestones of Middle Triassic
in Krížna Unit, Western Carpathians. Manuscript. Arch. Dep.
Geol. Paleont., Faculty Nat. Sci., Bratislava, 1—104 (in Slovak).
Tollmann A. 1976: Analyse des klassischen nordalpinen Mesozoi-
kums (Stratigraphie, Fauna und Fazies der Nördlichen Kalkal-
pen). Franz Deuticke, Wien, 1—580.
Tucker M.E. & Wright V.P. 1990: Carbonate sedimentology. Black-
well Sci. Publ., Oxford, London, Edinburgh, Boston, Mel-
bourne, 1—482.
Turanová L. & Turan J. 1993: Manometric analysis – a method to
quantitatively analyse Mg—Fe carbonates and their isomorphous
members. Acta Geol. Univ. Comen. 49, 17—22.
Vissher H., Van Houte M., Brugman W.A. & Poort R.J. 1994: Rejec-
tion of a Carnian (Late Triassic) “pluvial event” in Europe. Rev.
Palaeobot. Palynol. 83, 217—226.
Webb J. 1979: A reappraisal of the palaeoecology of conchostra-
cans (Crustacea: Brachiopoda). Neu. Jb. Geol. Paläont. 158,
259—275.
Wignall P.J. & Hallam A. 1992: Anoxia as a cause of the Permian/
Triassic mass extinction: Facies evidence from northern Italy
and the western United States. Palaeogeogr. Palaeoclimatol.
Palaeoecol. 93, 21—46.
Zenker J. 1834: Lingula keuperea Zenk. und Lingula calcaria
Zenk., zwei fossile Muschelarten aus Thüringen. Neu. Jb. Miner.
Geogn. Geol. Petref., 394—7.
Zonneveld J.-P., Beatty T.W. & Pemberton S.G. 2007: Lingulide
brachiopods and the trace fossil Lingulichnus from the Triassic
of Western Canada: implications for faunal recovery after the
end – Permian mass extinction. Palaios 22, 74—97.
è