GEOLOGICA CARPATHICA, 50, 2, BRATISLAVA, APRIL 1999
203211
FORAMINIFERA AND CALCAREOUS NANNOPLANKTON
ASSEMBLAGES FROM ?TITHONIAN-NEOCOMIAN CIESZYN
BEDS (SILESIAN UNIT, POLISH WESTERN CARPATHIANS)
ANDRZEJ SZYD£O and MA£GORZATA JUGOWIEC
Polish Geological Institute, Carpathian Branch, Skrzatów St. 1, 31-560 Kraków, Poland
(Manuscript received October 8, 1997; accepted in revised form March 24, 1998)
Abstract: The oldest micro- and nannofossils of the ?Tithonian Lower Cieszyn Shales and detrital Cieszyn Limestones
are dominated by redeposited calcareous benthic foraminifers and calcareous nannoplankton. These assemblages are
similar to those of epicontinental seas or large shelf areas. The younger micro- and nannofossil pelitic part of the
Berriasian Cieszyn Limestones and Valanginian Upper Cieszyn Shales are composed of low-diversity primitive agglu-
tinated foraminiferids and calcareous nannoplankton that resemble coeval faunas and floras of the Tethys seas. The
sequence of foraminiferal assemblages and the nature of the nannoplankton associations from the Cieszyn Beds
reflect the subsidence or collapse of the NE European Platform margin, disappearance of shallow areas with carbonate
sedimentation, and formation of a deep basin with turbidite sedimentation. These events may be related to the
Neocimmerian orogeny and the worldwide regression at the Tithonian/Berriasian boundary.
Key words: Tithonian-Neocomian, Polish Western Carpathians, Silesian Unit, Cieszyn Beds, paleoecology, calcareous
nannoplankton, Foraminifera.
Introduction
Preliminary analyses of foraminiferal and calcareous nanno-
plankton assemblages from the Silesian Series are presented.
These are part of the project dealing with the onset of the
subsidence of the Carpathian flysch basin. The biostrati-
graphical, paleoecological and paleogeographical studies re-
fer to the oldest sedimentary rocks (Cieszyn Beds) of the
Polish Western Carpathians. The microfossils of these de-
posits appear to be a valuable tool for paleoenvironment in-
terpretations and paleobiogeographical reconstructions. The
results of these investigations may be regarded as making
more precise and extending the former research about the
initial conditions for life and a sedimentation in the early fly-
sch basin (comp. Ksi¹¿kiewicz 1961, 1975; Olszewska
1984).
The study is based on samples collected from the Cieszyn-
Ustroñ area (Cisownica, Goleszów, Goleszów-Marglownia
Quarry), the vicinity of Bielsko-Bia³a (Lipnik Stream, Ka-
mienica Stream and Quarry, Jasienica Quarry) and neigh-
bourhood of the ¯ywiec tectonic window (So³a River, Radz-
iechowy Quarry), Fig. 1.
Geological setting
In the Polish Outer Carpathians, ?Tithonian non-flysch de-
posits and flysch-type Upper Tithonian-Lower Cretaceous
sediments belong to three tectonic units: Silesian, Subsile-
sian and Skole nappes. The oldest sediments of the Car-
pathians basin are more completely developed in the Silesian
Nappe of the Western Carpathians. The Silesian Nappe in
the study area consists of two independent tectonic units: the
Fig. 1. Location of the studied outcrops. Tectonic units in the Pol-
ish Western Carpathians (Geroch et al. 1967). Outcrops: 1Ci-
sownica, 2Goleszów; 3Goleszów-Marglownia Quarry, 4
Jasienica Quarry, 5Kamienica Stream and Quarry, 6Lipnik
Stream, 7Radziechowy Quarry, 8So³a River.
204 SZYDŁO
and JUGOWIEC
Cieszyn Unit and the Godula Unit (Bieda et al. 1963). The
first of these units consists of uppermost Jurassic (Tithonian)
and Lower Cretaceous sediments. Part of this sequence, the
so-called Cieszyn Beds belongs to the informal lithostrati-
graphical units: ?Tithonian Lower Cieszyn Shales (dark-
grayish marly shales), ?Upper Tithonian-Berriasian Cieszyn
Limestones (light-coloured, detrital and pelitic limestones)
and Valanginian-Hauterivian Upper Cieszyn Shales (dark-
grey, marly shales) (Fig. 2). The Cieszyn Beds as well as
the overlying Grodziszcze Beds (grey-bluish marls and
shales with rare sandy intercalations) have been studied for
micro- and nannofossils (Fig. 3).
The base of the Cieszyn Beds is uncertain because of the
tectonic decollement (Bieda et al. 1963) and the recently re-
vised Kimmeridgian-Tithonian boundary (Gradstein et al.
1995). Therefore, the analysis of the oldest part of these sed-
iments will be presented separately.
The characteristic, frequent and better preserved species
of foraminifers and calcareous nannoplankton are listed
(Fig. 3) and illustrated (Pls. I, II).
Results
Tithonian microfossils from Lower Cieszyn
Shales and detrital Cieszyn Limestones
Foraminifera
In Goleszów section (cf. Szyd³o 1997) and
from Marglownia Quarry, the oldest samples (i.e.
?Upper Tithonian age) yield mainly calcareous
benthos: Geinitzinita wolinensis Bielecka &
Po¿aryski, few Lenticulina: L. ex gr. münsteri
(Roemer), L. cf. ambanjabensis Epistalié & Sigal,
Lenticulina vistulae elongata Bielecka &
Po¿aryski, numerous taxa of Planuria sp.: P.
cordiformis (Terquem), P. crepidularis Roemer,
P. multicostata Kusnetzova, P. cf. poljenovae
Kusnetzova, Vaginulinopsis embaensis (Fursenko
& Polenova), Marginulinopsis robusta (Reuss),
M. striatocostata (Reuss), Tristix temiricra
(Dain), Vaginulina kochii Roemer and Trocholina
solecensis Bielecka & Po¿aryski. In addition oth-
er badly preserved nodosariids (e.g. Lenticulina
spp., Lingulina spp., Frondicularia spp., Cithari-
na spp.) are noted. Moreover, from Marglownia
Quarry common taxa of Polymorphinidae: Eogut-
tulina liassica (Strickland), Guttulina multistriata
Bielecka are reported. Agglutinated foraminifers,
including Belorusiella wolinensis Bielecka,
Palaeogaudryina cf. taurica (Gorbachik),
Palaeogaudryina varsoviensis (Bielecka &
Po¿aryski) as well as radiolarians, diatoms and
fragments of ostracods, bryozoans and corals
have also been found in these deposits.
At the top of the Lower Cieszyn Shales, just
below the Cieszyn Limestones (Cisownica) the
uppermost Tithonian associations are noted. The
first occurrence of Trocholina group: T. alpina
(Leupold), T. solecensis Bielecka & Po¿aryski
Fig. 2. The Cieszyn Beds in the Polish Western Carpathians (Cieszyn Unit)
based on Geological atlas of Poland 1962; Nowak 1973.
and Neotrocholina molesta (Gorbachik), and diverse Lenti-
culina: L. infravolgensis Fursenko & Polenova, L. münsteri
(Roemer), L. ponderosa Mjatliuk, L. cf. vistulae Bielecka &
Po¿aryski are observed. In addition, Marginulinopsis betten-
staedti (Bartenstein & Brand), M. striatocostata (Reuss), Sa-
racenaria alata-angularis (Franke), Paalzowella feifeli
(Paalzow), Spirillina minima Schacko have been distin-
guished (Szyd³o 1997). The above foraminiferids are charac-
teristic for the detrital Cieszyn Limestones (Geroch 1966).
Calcareous nannoplankton
The samples studied from the Lower Cieszyn Shales con-
tain poor nannoplankton assemblages. Most of the samples
are dominated by species of Watznaueria Reinhardt (which
are characteristic of poorly preserved assemblages) as well
as by Ellipsagelosphaera Noël, which constitute about 95 %
of the assemblages. Identified species are as follow: W. bar-
nesae (Black) Perch-Nielsen, W. biporta Bukry, E. britanni-
ca (Stradner) Perch-Nielsen, E. fossacincta Black, E. lucassi
FORAMINIFERA AND CALCAREOUS NANNOPLANKTON ASSEMBLAGES 205
Noël. At the boundary with the Cieszyn Limestones other
species (up to 5 in the sample) have been encountered. They
are: Conusphaera cf. mexicana Trejo, Cyclagelosphaera de-
flandrei (Manivit), C. margerelii Noël, Diazomatolithus leh-
manii Noël, Microstaurus chiastus (Worsley) Grün, Polycos-
tella cf. beckmannii Thierstein, Zeugrhabdotus embergei
(Noël) Perch-Nielsen.
Samples collected from the uppermost part of the Lower
Cieszyn Shales probably represent at least the top of the
Zeugrhabdotus embergeri Zone of Worsley (1971). The nan-
noplankton assemblages of this zone, dominated by Ellip-
sagelosphaera. FO of C. mexicana, P. beckmannii, indicate
the top of this zone (Wind 1978; Hamilton 1982), i.e. the up-
permost Early Tithonian. However, according to studies by
Bralower et al. (1989) from land sections of south-western
Europe, FO of P. beckmannii indicates at least the lower part
of the Upper Tithonian.
The oldest assemblages described from the non-flysch
marly Lower Cieszyn Shales are dominated by calcareous
benthic foraminifers and scarce nannoplankton. Assemblag-
es with common trocholinas also resemble the microfauna of
detrital Cieszyn Limestones (cf. Geroch 1966). Samples
from Radziechowy and Jasienica limestones quarries are de-
void of microfossils. The accompanying radiolarians as well
as the style of deposition of these sediments (S³omka 1986)
suggest that the shallow-water calcareous foraminifers
(mainly nodosariids and trocholinas) and nannoplankton
taxa (e.g. Conusphaera mexicana, Thierstein 1976) have
been redeposited.
Berriasian and Valanginian microfossils from Cieszyn
Limestones (pelitic part) and Upper Cieszyn Shales (lower part)
Foraminifera
In the Kamienica Quarry, improverished and badly pre-
served assemblages are found, in shally intercalation of the
Cieszyn Limestones. They consist of numerous primitive ag-
glutinated forms such as: Rhizammina indivisa Brady, Hy-
perammina gaultina Dam, Ammodiscus sp., Glomospira sp.,
Pseudoreophax cisovnicensis Geroch, Trochammina quin-
queloba Geroch and scarce calcareous benthic taxa (e.g. Tro-
cholina paucigranulata Moulladae). Forms belonging to
Nodosariidae (Lenticulina) and Involutinidae (Trocholina,
Neotrocholina) have corroded tests. A few radiolarians and
ostracods were also observed.
Calcareous nannoplankton
The calcareous nannofossil assemblages of the Cieszyn
Limestones are similar to those from the Lower Cieszyn
Shales. However, Nannoconus spp. Kamptner and Speetonia
colligata Black have been found in one sample. The exam-
ined material from the Upper Cieszyn Shales has been de-
void of calcareous nannoplankton. Nevertheless, occurrence
of Nannoconus spp. and S. colligata may suggest at least the
Early Berriasian age of the studied samples.
Low-diversity and poor foraminiferal assemblages are
characteristic of the Berriasian-Valanginian Cieszyn Lime-
Fig. 3. Table of proposed distribution of some taxa of foraminifers and calcareous nannoplankton.
206 SZYDŁO and JUGOWIEC
stones (pelitic part) and Upper Cieszyn Shales (without their
uppermost part; compare with Geroch 1966). These assem-
blages are composed mainly of agglutinated taxa belonging
to: Ammodiscidae, Ataxophragmiidae, Astrorhizidae and
may be consider as autochthonous fauna. These microfossils
represent the slope of a deep flysch basin (cf. Ksi¹¿kiewicz
1961, 1975; Olszewska 1984). Moreover, the sedimentologi-
cal data confirm this opinion (S³omka 1986; Malik 1986,
1994).
Uppermost Valanginian-Hauterivian microfossils from
Upper Cieszyn Shales (uppermost part) and Grodziszcze
Beds (lower part)
Foraminifera
Microfauna of the Upper Cieszyn Shales (Lipnik Stream,
So³a River) and the lower part of the Grodziszcze Beds (Lip-
nik Stream) includes mainly Ataxophragmiidae and scarce
Nodosariidae (mainly Lenticulina with corroded tests). To
the former belong Praedorothia hauteriviana (Moullade)
with calcareous agglutinated test and arenaceous species:
Pseudoreophax cisovnicensis Geroch and Verneuilinoides
neocomiensis (Mjatliuk) which are characteristic of the Up-
per Cieszyn Shales (upper part) and the Grodziszcze Beds
(cf. Geroch 1966). Other forms i.e. Falsogaudryinella teal-
byensis (Bartenstein), Ammobaculoides carpathicus Geroch
also occur in the Grodziszcze Beds samples.
Calcareous nannoplankton
The similarity of specific compositon of nannofossil as-
semblages between the Grodziszcze Beds and underlying
beds (Lower Cieszyn Shales, Cieszyn Limestones) have
been observed. Moreover, the presence of Haqius circumra-
diatus (Stover) Roth, Lithraphidites sp. Deflandre, and Tu-
bodiscus cf. verenae Thierstein in the Grodziszcze Beds
samples have been noted. According to Crux (1982) the
range of H. circumradiatus is from the Hauterivian to the
Campanian, or according to Mutterlose (1991) from the
Kimmeridgian to the Barremian in Tethys. The presence of
T. cf. verenae probably suggest a Valanginian age of the in-
vestigated samples (Mutterlose 1989).
The microfauna indicates outer shelf and slope environ-
ments for the flysch basin (cf. Malik & Olszewska 1984).
Mixed foraminiferal assemblages with poorly preserved and
low diversified nannoplankton associations may be connect-
ed with tectonic activity of the flysch basin (Haq 1973;
Ksi¹¿kiewicz 1975; S³omka 1986), caused by sea level fluc-
tuations (Haq 1973; Cooper 1977; Leszczyñski & Malik
1996) or climate changes (Haq 1973).
Conclusion
The microfauna characteristic of the northen margins of
Tethys dominating the Tithonian sediments was replaced by
Neocomian flysch taxa, which represent the slope environ-
ment of the Cieszyn Basin.
The Tithonian foraminiferal assemblages containing high-
ly diversified calcareous benthos of the families: Nodosari-
idae (Geinitzinita, Lenticulina, Marginulinopsis, Planularia,
Saracenaria, Vaginulina, Vaginulinopsis), Involutinidae
(Trocholina, Neotrocholina, Paalzowella), Polymorphinidae
(Eoguttulina, Guttulina) and minor agglutinated foramin-
iferids belong to Ataxophragmiidae family (Paleogaudryi-
na, Belorusiella) may be correlated with the shelf assem-
blages of Gordon (1970).
Certain species of Watznaueria and Ellipsagelosphaera
are well preserved and dominate calcareous nannoplankton
assemblages. The other identified taxa are relatively poorly
preserved. C. mexicana and Nannoconus spp. most probably
preferred the epicontinental seas and large shelf areas (cf.
Thierstein 1976).
Redeposited foraminiferids of the Lower Cieszyn Shales
and Cieszyn Limestones (Geroch 1966; Nowak 1976;
Geroch & Olszewska 1990), the nature of calcareous nanno-
plankton (Thierstein 1976; Perch-Nielsen 1979), associated
with the presence of radiolarians (Ksi¹¿kiewicz 1975) con-
firm the opinion that carbonate material was transported by
turbidites into a deeper environment (at least an upper bathy-
al zone) from the northern platform or reef margins of Sile-
sian (Cieszyn) Basin (Geological Atlas of Poland 1962; At-
las of paleotransport 1976, S³omka 1986; Michalík & Soták
1990). According to Sliter (1980), shallow-water exchange
between the Boreal and Tethys realms was initiated at about
150 Ma (i.e. during Malm).
At the Jurassic/Cretaceous boundary, the first agglutinated
foraminiferal population appeared in the Cieszyn Lime-
stones (with Trochammina sp.) as microfauna that started to
colonize the new created flysch environment in the Outer
Polish Carpathians Basin (Olszewska 1982). This trend is
represented by autochthonous assemblage with Pseu-
doreophax cisovnicensis which occurred in the Berriasian-
Valanginian Cieszyn Limestones and the Upper Cieszyn
Shales (without the uppermost part).
Poor and badly preserved assemblages with Pseu-
doreophax cisovnicensis resemble the Recurvoides Asso-
ciation of Haig (1979) and represent the slope of the flysch
basin (bathyal zone) close (pelitic Cieszyn Limestones) or
below (Upper Cieszyn Shales) the local CCD level.
Plate I: Fig. l. Palaeogaudryina varsoviensis (Bielecka &
Po¿aryski). Fig. 2. Pseudoreophax cisovnicensis Geroch. Fig. 3.
Verneuilinoides neocomiensis (Mjatliuk). Fig. 4. Geinitzinita
wolinensis Bielecka. Fig. 5. Lingulina sp. Fig. 6. Lenticulina vis-
tulae elongata Bielecka & Po¿aryski. Fig. 7. Lenticulina ex gr.
münsteri (Roemer). Figs. 810. Lenticulina sp. Fig. 11. Planularia
cordiformis (Terquem). Fig. 12. Planularia multicostata (Kusnet-
zova). Fig. 13. Planularia crepidularis Roemer. Fig. 14. Planular-
ia uilensis Kusnetzova. Fig. 15. Vaginulina kochii Roemere. Fig.
16. Marginulinopsis cf. bettenstaedti (Bartenstein & Brand). Fig.
17. Marginulinopsis striatocostata (Reuss). Fig. 18. Eoguttulina
liassica (Stricland). Fig. 19. Guttulina multicostata Bielecka. Fig.
20. Vaginulinopsis embaensis Fursenko & Polenova. Fig. 21. Tris-
tix acutangulus (Reuss). Fig. 22. Trocholina solecensis Bielecka &
Po¿aryski. Figs. 2324. Neotrocholina molesta Gorbachik. Fig.
25. Trocholina alpina (Leupold). Fig. 26. Paalzowella feifeli
(Paalzow). Length of bars: 0.1 mm.
v
PLATE I 207
208 PLATE II
FORAMINIFERA AND CALCAREOUS NANNOPLANKTON ASSEMBLAGES 209
The low-diversity agglutinated microfauna (mainly with P.
cisovnicensis) may be correlated with the first oxygen mini-
mum episode in the Cieszyn Basin (comp. Szyd³o 1997).
The foraminiferal morphotypes of these assemblage contains
mainly small, elongated, planispiral-flattened and cylindrical
forms with high surface areato volume ratios. This fact
may be related to the extent of anoxia (Bernhard 1986).
Probably, during this time (i.e. earliest Cretaceous) the sup-
ply of terrigenous material to the vast flysch basin increased.
Rapid rise of the CCD, related to the restricted conditions
(e.g. stagnation of the bottom water) caused elimination of
benthic life and, consequently, occurrence of very impover-
ished foraminiferal associations (Butt 1977; Kaminski et al.
1995; Holbourn & Kaminski 1997).
The uppermost Valanginian-Hauterivian Upper Cieszyn
Shales and Grodziszcze Beds yielded both arenaceous and
rare calcareous foraminifers which the resemble Mars-
sonella/Recurvoides associations of Haig (1979). The spe-
cific composition of these foraminiferal assemblages with
the accompanying scarce calcareous nannoplankton most
probably reflect a transgression-regression cycle in the fly-
sch basin (Haq 1973; Leszczyñski & Malik 1996) connected,
in turn, with the short duration of tectonic activity of the
northen margins of the basin at the end of the Valanginian
(S³omka 1986).
The specific composition of nannofossil assemblages
(poor, low-diversity, badly preserved) has also been ob-
served in Neocomian sediments (from the pelitic Cieszyn
Limestones to the Grodziszcze Beds).
Widespread orogenic events and regression episodes are
followed by reduction in the quantity of calcareous nannof-
lora (Haq 1973).
The sequence of foraminiferal assemblages and the nature
of the nannoplankton association from the Cieszyn Beds
reflect the subsidence or collapse of the NE European mar-
gin of the platform, the disappearance of areas with shallow
carbonate sedimentation and formation of a deep basin with
flysch sedimentation.
The geotectonic desintegration of the carbonate platform
system preceded formation of the Carpathian deep flysch ba-
sin (Leszczyñski & Malik 1996). Therefore, the Neocimme-
rian orogeny (Nowak 1976) and the worldwide regression at
the Tithonian/Berriasian boundary (Zeiss 1983) may be re-
sponsible for this event.
Foraminifers and some taxa of calcareous nannofossils,
reported from the ?Tithonian Lower Cieszyn Shales are com-
parable to those of epicontinental seas or large shelf areas,
for example the European Platform (cf. Bielecka &
Po¿aryski 1954; Bielecka & Geroch 1974; Bielecka 1975;
Thierstein 1976; Kuznietzova & Gorbachik 1985). Their ori-
gin may be related to the sedimentation area of the Pavlov
platform (Hanzliková 1965a: the youngest part of the Klent-
nice Beds; Eliá & Eliáova 1986). The presence of certain
foraminiferids (e.g. trocholinas) with fragments of macro-
fauna (e.g. bryozoans, ostracods, corals) in the Lower
Cieszyn Shales may represent part of the eroded material
from the nearby Inwa³d klippe (Ksi¹¿kiewicz 1971; S³omka
1986).
Moreover, species of Watznaueria and Ellipsagelosphaera
and Z. embergeri have consistent occurrences in both the
Tethyan and Boreal realms. Thus, these species represent an
eurytropic flora (Muterlose 1992).
The microfauna described from the Neocomian deposits in
the Cieszyn flysch basin (northern margin of Tethys) sug-
gests that the sedimentation conditions were similar to those
of the Slovak Western Carpathians and deeper (scarce Epis-
tommina) than those in the Western Mediterranean region
(Borza et al. 1995). However, bathimetrically these condi-
tions were similar to those calculated for the Slovak Car-
pathians (Hanzliková 1956a,b; Andrusov 1959), Romanian
Carpathians (Neagu 1962), Eastern Alps (Decker & Rögl
1988) and Betic Mountains (Kuhnt 1995), and shallower
than those from the North Atlantic (Sliter 1980). Certain
cosmopolitan species e.g. Rhizammina indivisa, Hyperam-
mina gaultina, Pseudoreophax cisovnicensis, Trochammina
quinqueloba, have been noted in the coeval sediments of
the Indian Ocean (Holbourn & Kaminski 1997).
The Early Cretaceous nannofossils: C. mexicana, Nanno-
conus spp., S. colligata, P. beckmannii have been restricted
to the tropical and subtropical paleolatitudes. However, C.
mexicana and Nannoconus spp. are characteristic of the
Tethyan paleobiogeoprovince. Nevertheless, they are very
scarce or absent in the Pacific realm (cf. Thierstein 1976;
Perch-Nielsen 1979; Mutterlose 1992).
Mixed microfossils from different environments have
been distinguished from Tithonian sediments (Lower
Cieszyn Shales, detrital Cieszyn Limestones) and uppermost
Valanginian-Hauterivian deposits (uppermost part of the Up-
per Cieszyn Shales, Grodziszcze Beds), consequently these
sediments may be interpreted as resedimented (Olszewska
1983; S³omka 1986; Leszczyñski & Malik 1996).
Acknowledgements: Special thanks are offered to P. Ne-
scieruk M. Sc. for supplying some of the samples for this
study. Thanks are extended to Dr. B. Olszewska, Dr. A.M.
Gasiñski, Dr. J. Soták for their critical and helpful remarks
on the manuscript.
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