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Department of Geology and Paleontology, Faculty of Sciences, Mlynská dolina, 842 15 Bratislava, Slovak Republic

(Manuscript received February 6, 1997;  accepted in revised form June 24, 1997)


: The paper concerns all Eocene anascan Bryozoa known from Slovakia. New bryozoan associations from

Štrba and Poluvsie are studied and described. A total of 10 taxa, newly found in Slovakia are described, the remainder
are listed.

Key words: 

Western Carpathians, Eocene, Bryozoa, Anasca, systematics.


This paper concerns anascan Bryozoa from Eocene sedi-
ments, in the Slovak part of the Western Carpathians (Fig. 1).
The associated fieldwork (done within the last two years)
was aimed at finding new bryozoan localities not described
in the previous paper of Zágoršek (1994). Two localities —
Štrba and Poluvsie yielded bryozoan associations.

The bryozoan zoaria mostly occur in well lithified marls.

Several techniques have been used to have specimens with
observable surface. Most of the techniques are aggressive,
which caused bad preservation of the fossils. However, most
of the specific features have been preserved and could be ob-
served. Some of the found specimens cannot be listed under
any known species, but due to the bad preservation, no new
species could be erected.

Most of the Eocene anascan Bryozoa from the Slovak part

of the Western Carpathians have been published already
(Zágoršek 1996a,b). The remainder are described here.

The holotypes and paratypes of Reuss species as well as of

Stoliczka species have been studied in the Vienna Natural
History Museum in 1996. The original material has been
compared with the material from Slovakia. All observations
are commented on, in the “Remarks” within the description
of the species.

Description of the localities

The Eocene sediments of the Western Carpathians can be

divided into four formations (Gross et al. 1984). The studied
bryozoans are from the basal Borové Formation (Východná,
Partizánska  upča, Hybica and Štrba) and from the Zuberec
Formation (Rajecké Teplice and Poluvsie).

The lithological content of the Borové Formation in the

Liptov Basin is rather varied (Gross & Köhler 1980). It con-
sists of fine-grained calcarenites (Východná), or of
yellowish-brown organodetrital clayey limestone (Partizáns-
ka  upča). Within the formation, there are a thin-bedded,
pale-grey to brown bryozoan limestones (Hybica and Štrba).

The locality at Východná (Fig. 2) is situated east of the

village, in a road-cut near the railway station. The fossils
(nummulites, bryozoans and rare molluscs) occur in an orga-
nodetrital limestone. Among them are Nummulites garnieri
Boussac and Nchavannesi de la Harpe, from which an
Eocene age has been determined (Gross & Köhler 1980). The
locality probably represents a shallow marine environment.
This can be confirmed by the occurrence of the bryozoan
genera Lacrimula and Lunulites which lived only in shallow
marine conditions in the Eocene (Braga & Munari 1972).

The locality at Partizánska  upča (Fig. 3) is in an aban-

doned quarry lying about 1.6 km south of the village. Fora-
miniferal tests (Discocyclina prattii (Michelin 1846) and
Nummulites chavannesi 

de la Harpe) dominate over other al-

lochems in the sediment. Nummulites chavannesi determines
the Eocene age (Gross & Köhler 1980). As in the locality Vý-
chodná, the large quantity of large forams as well as the pres-
ence of Lacrimula point to a shallow marine environment.

The locality called Hybica (see Fig. 2) is situated on the

right bank of the Hybica River, about 3.4 km north of the vil-
lage of Východná. The outcrop represents the upper part of a
biogenic (mostly algal) buildup. Bryozoan marl occurs as a
lensoid body, probably formed by a slump, in the claystone
of the Huty Formation. According to Zágoršek (1996) the
original sedimentation area of the bryozoan marls could be a
shelf margin with cool water due to the influence of up-
welling. Later, marls with bryozoans probably slumped to-
ward the basin and into deeper sediments.

Fig. 1.

 Sketch of the geographical position of the described locali-

ties. 1 — Východná, 2 — Hybica, 3 — Partizánska  upča, 4 —
Štrba, 5 — Rajecké Teplice, 6 — Poluvsie.

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402                                                                                              ZÁGORŠEK

The Štrba locality lies in the abandoned railway cut, about

100 m north of Kolombiarok Hill (899.6 m) and about 800 m
east of the road connecting the villages of Tatranská Štrba
and Štrba (see Fig. 4). This locality represents the best exam-
ple of the bryozoan marl facies. Probably, it is a large lensoi-

dal, slump body originated in a similar environment to the
sediments at Hybica. However, abundant large shallow ma-
rine molluscs have been found here. In the upper part of the
section, Nummulites boullei de la Harpe and Operculina al-

Douvillé, have been found. These prove an Eocene age

(Köhler, personal comm, 1995) of these sediments.

The  Zuberec Formation represents flysch type sediments.

These are calcareous sandstones interbedded with clay-
stones and rarely with embedded limestones. The Bryozoa
occur as grains within the sandstones.

At  Rajecké Teplice (see Fig. 5), bryozoans occur in a

medium-grained calcarenite, exposed above the Bystrička
creek, south of the village.

The bryozoans from Poluvsie (Fig. 6) were found in rocks

collected in the field around the hill called Diel (473.4 m).
The rock is similar to that at Rajecké Teplice, an yellowish to
brownish medium-grained calcarenite.

The depositional environments of these localities cannot

be suggested due to the small amount of remains of bryozo-
ans as well as other fossils.


The anascan Bryozoa have a zooecial frontal surface

formed by an uncalcified frontal membrane. Calcified cryp-
tocyst usually lies beneath these membrane, and can be short
(Pseudomalacostega and Cribrimorpha) or extend forward
towards the aperture (Coilostega and Pseudostega). The
space between membrane and cryptocyst forms part of the
hydrostatic system used in tentacle protrusion. The system
used parietal muscles which depress the frontal membrane
and reduce the space between it and the cryptocyst. These
parietal muscles are located in various places, mostly being

Fig. 2.

 Sketch of the geographical position of Východná (1) and Hy-

bica (2) localities.

Fig. 3.

 Sketch of the geographical position of Partizánska  upča

locality (arrow).

Fig. 4.

 Sketch of the geographical position of Štrba locality (arrow).

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The proximal part of the zooecia, not covered by the frontal

membrane, may form a calcified frontal wall called the gym-

Interzooecial avicularia and/or vibracula and ovicells are

generally present.


All of the anascan Bryozoa found have been listed in Table

1 with references to their last description. The higher taxa are
listed according to Gordon (1986).

Class Gymnolaemata Allman 1896

Order Cheilostomata Busk 1852

Suborder Anasca Levinsen 1909

Division Pseudomalacostega J.& L. d’Hondt 1977

Family Calloporidae Norman 1903

Genus Amphiblestrum Gray1848

Amphiblestrum appendiculata 

(Reuss 1848)

(Pl. I: Fig. 1)

1848 Cellepora appendiculata sp.n., Reuss p. 96, Pl. 11, Fig. 22
1963 Ramphonotus appendiculatus (Reuss 1848), Braga p. 23
1974 Ramphonotus appendiculata (Reuss 1848), David & Pouyet p.
           108, Pl. 1, Figs. 2, 6
1994 Rhamphonotus monopora Reuss, Zágoršek tab. 1
1996a Ramphonotus sp., Zágoršek p. 125, Pl. 2, Fig. 2


Zoaria encrusting. Zooecia membraniporiform,

oval to triangular, with very short cryptocyst. Opesia triangu-
lar. A shallow, narrow furrow separates neighboring zooecia.
No gymnocyst. Mural rim narrow, same width around all zoo-
ecium, smooth. Vibracula rare, usually situated among three or
four autozooecia, small, tube-like, with circular orifice.

The described specimens are identical in shape of

the zooecia and shape and location of the vibracula with the
holotype deposited in the Natural History Museum in Vien-
na. However the holotype mural rim is slightly ribbed.

Gordon (1984) considered Ramphonotus to be a junior

synonym of Amphiblestrum. Small gymnocyst, development
of the cryptocyst and position of the avicularia characterized
genus Amphiblestrum.  Presence of these features in the de-
scribed specimens allow us to list this species under the ge-
nus Amphiblestrum.

Štrba locality, probably also in the Poluvsie

and the Partizánska  upča localities.

Division Coilostega Levinsen 1902

Family Lunulitidae Lagaaij 1952

Genus Lunulites Lamarck 1816

Lunulites quadrata 

(Reuss 1848)

(Pl. II: Fig. 5)

1848 Cellepora quadrata sp.n., Reuss p. 95, Pl. 11, Fig. 17
1869 Lunulites quadrata (Reuss 1848), Reuss p. 278, Pl. 28, Fig. 16
1963 Lunulites quadrata (Reuss 1848), Braga p. 25, Pl. 2, Fig. 8
1980 Lunulites quadrata (Reuss 1848), Braga p. 47, Fig. 37.
1988 Lunulites quadrata (Reuss 1848), Braga & Barbin p. 518.

Fig. 5.

 Sketch of the geographical position of Rajecké Teplice lo-

cality (arrow).

Fig. 6.

 Sketch of the geographical position of Poluvsie locality (arrow).

restricted to the proximal margin of the orifice or passing in
pairs through openings in the cryptocyst called opesiules.

The frontal membrane is surrounded by an elevated, calcified

margin called the mural rim. In the Cribrimorpha, the membra-
nous frontal is covered by a costal frontal shield (pericyst).

The true opening, where tentacles are protruded is called

the orifice. However, when the membranous frontal wall does
not fossilize, the preserved opening is called the opesium.

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Zoaria free, conical, cupuliform disk in shape.

Preserved are about one third of the whole colony. Rectan-
gular zooecia arranged in radial rows. Orifice oval to circu-
lar. Mural rim narrow. The cryptocyst developed. Ovicell
endozooecial, have not been observed. Avicularia as well as
vibracula present small, oval.

Only a fragment of a zoarium has been found.

However, the shape and morphology of the zooecia allow us
to determine the most probable species, but many of the spe-
cific features have not been preserved (mainly the ovicells).
Lunulites quadrata 

(Reuss 1848) is the most common spe-

cies of the genus in other Eocene sediments.


Family Cothurnicellidae Bassler 1935

Genus Chlidoniopsis Harmer 1957

Chlidoniopsis vindobonensis 

(Reuss 1848)

(Pl. I: Figs. 2-3)

1848 Crisidia vindobonensis sp.n., Reuss p. 54, Pl. 7, Fig. 25
1869 Unicrisia tenerrima sp.n., Reuss p. 279, Pl. 34, Fig. 7
1891 ?Catenaria tenerrima (Reuss), Waters p. 5, Pl. 1, Fig. 11
1963 Catenicella tenerrima (Reuss), Braga p. 40, Pl. 4, Fig. 2
1980 Chlidoniopsis vindobonensis (Reuss 1848), Braga p. 49,
           Figs. 40, 41
1988 Chlidoniopsis tenerrima (Reuss), Braga & Barbin
           p. 519, Pl. 6, Figs. 5, 8


Zoaria erect, not articulated. Zooecia drop-like,

narrow, very long with enormously elongated proximal part —
canda. The canda is as long as the rest of zooecium, sometimes
even longer. Orifice terminal, with wide sinus on the proximal
border. No avicularia or ovicells.

The described specimens are identical with holo-

type of Crisidia vindobonensis Reuss 1848 as well as of the
holotype of Unicrisia tenerrima Reuss 1869 deposited in the
Natural History Museum in Vienna. Because Crisidia vindo-

Reuss 1848 is the first description of this species it

should be used instead of Chlidoniopsis tenerrima (Reuss).
Braga (pers. comm, 1997) argues, that Reuss (1869) himself
changed the name vindobonensis to tenerrima, because the
species does not occur in the Vienna Basin but in Val di Lon-
te in Italy. However the name of species cannot be changed
even by the author himself (according to Intern. Code of No-

mencl.). So the occurrence of the species is not a reason for
changing the species name, neither is it incorrect.


Division Pseudostega Levinsen 1909

Family Cellariidae Hincks 1880

Genus Cellaria Ellis & Solander 1786

Cellaria reussi 

d’Orbigny 1851

(Pl. I: Figs. 4, 6)

1869 Salicornaria Reussi d’Orbigny 1851, Reuss p. 261, Pl. 29, Fig. 5
1963 Cellaria reussi d’Orbigny 1851, Braga p. 26
1980 Cellaria reussi d’Orbigny 1851, Braga p. 50, Fig. 43
1988 Cellaria reussi d’Orbigny 1851, Braga & Barbin p. 519, Pl. 6,
           Fig. 7


Zoarium articulated, with cylindrical segments

(internodes). Zooecia in four longitudinal rows, alternating
with each other. Zooecia drop-like in shape, proximally taper-
ing. Mural rim narrow, smooth. Cryptocyst concave, exten-
sive, smooth. Orifice semilunar with enlarged proximo-lateral
corners for parietal muscles. No avicularia. Ovicell endozooe-
cial, small, have not been observed.

Each zoarium can yield huge numbers of intern-

odes, which is the reason for the large abundance of Cellaria
fragments in the fossil record.

This is a common species at Štrba locality and

is rarely found also at Hybica.

Family Scrupocellariidae Levinsen 1909
Genus Scrupocellaria van Beneden 1845

Scrupocellaria brendolensis 

Waters 1891

(Pl. I: Fig. 7; Pl. II: Figs. 3–4)

1891 Scrupocellaria brendolensis sp.n., Waters p. 7, Pl. 1, Figs. 14, 15
1975 Scrupocellaria brendolensis Waters 1891, Braga p. 146, Pl. 2,
          Figs. 1, 2
1988 Scrupocellaria brendolensis Waters 1891, Braga & Barbin p.
         520, Pl. 6, Figs. 2–3


Zoaria erect unilamellar (cellariiform), with two

rows of zooecia. Zooecia with opesia as long as one half of the
zooecial length. On the proximal margin of opesia, there is a
triangular to oval avicularium with pivotal bar. A small, circu-
lar vibraculum is attached near the distal margin of the opesia.
On the dorsal surface, each zooecium has a vibraculum.

The specimens described by Braga & Barbin

(1988) from Bressana have narrower zoaria and a small fur-
row separating adjacent zooecia on the dorsal surface of the
zoarium. These features are not described by Waters (1891),
however it could be explained by within species variation.

Two specimens found in Štrba have no frontal avicularia.

Scrupocellaria brendolensis 

Waters 1891 differs from

Scrupocellaria elliptica 

(Reuss 1848) mainly in having avic-

ularia on the frontal surface of the zoarium. However the
avicularia could be missed due to the bad presevation of the
specimens. So these two specimens more probably belong
to Scrupocellaria brendolensis Waters 1891 than to Scrupo-

Plate I:

 Fig. 1. Amphiblestrum appendiculata (Reuss 1848) —

Štrba locality, SEM BS 300, photo by I. Holický. Figs. 2–3. Chli-
doniopsis vindobonensis 

(Reuss 1848) — Štrba locality, SEM BS

300, photo by I. Holický. Fig. 4Cellaria reussi d’Orbigny 1851 —
Štrba locality, SEM BS 300, photo by I. Holický. Detail of the zoo-
ecium showing orifice with enlarged proximo-lateral corners and
cryptocyst. Fig. 5. Puellina (Cribrilaria) scripta (Reuss 1848) —
Hybica locality, photo by the author. Fig. 6. Cellaria reussi
d’Orbigny 1851 — Hybica locality, SEM Jeol, photo by J. Kulich.
Fig. 7.

 Scrupocellaria brendolensis Waters 1891 — Štrba locality,

SEM BS 300, photo by I. Holický. The specimens with no frontal
avicularia. The bad preservation does not allow us to judge, if it is
Scrupocellaria brendolensis Waters 1891 or Scrupocellaria ellip-

Reuss 1848. Scale bars represent 0.1 mm.

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406                                                                                               PLATE II

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cellaria elliptica 

(Reuss 1848), which is the Miocene spe-

cies (Schmid 1989).

Štrba locality

Division Cribrimorpha Lang1916
Family Cribrilinidae Hincks 1880

Genus Cribrilaria Canu & Bassler 1929

Cribrilaria radiata 

(Moll 1803)

(Pl. II: Figs. 6–7)

1920 Puellina radiata Moll, Canu & Bassler p. 294, Fig. 84/G-J,
           Pl. 41, Figs. 14–18
1974 Cribrilaria radiata (Moll), David & Pouyet p. 136
1980 Cribrilaria radiata (Moll), Braga p. 51, Fig. 50a
1988 Cribrilaria radiata (Moll), Braga & Barbin p. 521


Zoaria encrusting. Zooecia oval with semilu-

nar aperture. The width of the zooecium ranges from 0.146
mm to 0.261 mm and length is from 0.232 mm to 0.322 mm.
Frontal wall consists of 6–8 pair of costae with very narrow
lateral costal fusions. Median lamella not developed. Aper-
ture semilunar. Oral spines sometimes missing, sometimes
developed. Avicularium large, intrazooecial, with long ros-
trum, without pivotal bar. Hyperstomial ovicell with smooth
frontal surface.

According to David & Pouyet (1974) Cellepora


Reuss 1848 and Cellepora megacephala Reuss 1848

are junior synonyms of Eschara radiata Moll 1803. However
Schmid (1989) pointed out that Cellepora scripta Reuss
1848 (and Cellepora megacephala Reuss 1848 as a junior
synonym) is a different species — thus named Puellina
(Cribrilaria) scripta 

Reuss 1848. Studying the holotype of

Reuss material and reading the description of Cribrilaria ra-

(Moll) by David & Pouyet (1974) I agree with

Schmid’s (1989) opinion and do not list Cellepora scripta
Reuss 1848 and Cellepora megacephala Reuss 1848 among
the synonyms of Cribrilaria radiata (Moll).

A common species at Hybica and Štrba localities

Cribrilaria haueri 

(Reuss 1848)

(Pl. II: Fig. 1)

1848 Cellaria haueri sp. n.Reuss p. 63, Pl. 8, Fig. 9
1848 Eschara crenatimargo sp.n.Reuss p. 72, Pl. 8, Fig. 38
1869 Eschara haueri (Reuss 1848), Reuss p. 271, Pl. 32, Figs. 14–16
1963 Cribrilaria haueri (Reuss), Braga p. 29, Pl. 3, Figs. 9–10
1988 Cribrilaria haueri (Reuss), Braga & Brabin p. 521, Pl. 7, Fig. 1


Zoaria adeoniform, bifoliate, with 3 to 8 rows

of zooecia. Zooecia elongate, two times longer than wide.

Frontal wall consists of about 10 pairs of smooth costae.
Lateral costal fusions are developed, producing 5 to 6 pores
between each two costae. Median lamella not developed.
Gymnocyst very short or not developed. Aperture semicir-
cular to oval with apertural spines. Ovicell hyperstomial,
oval, little elongated, with smooth surface. No avicularia.

According to Braga (1991) Eschara crenatimar-


Reuss 1848 is the junior synonym of this species. The

described specimens are identical with the holotype deposit-
ed in the Natural History Museum in Vienna.

Hybica locality

Genus Puellina Jullien 1886

Puellina (Cribrilaria) scripta 

(Reuss 1848)

(Pl. I: Fig. 5)

1848 Cellepora scripta sp.n., Reuss p. 82, Pl. 9, Fig. 28
1848 Cellepora megacephala sp.n., Reuss p. 83, Pl. 10, Fig. 5
1989 Puellina (Cribrilaria) scripta (Reuss 1848), Schmid p. 26,
         Pl. 6, Fig. 10 (cum. syn.)


Zoaria encrusting. Zooecia oval to circular. The

average width of the zooecium is 0.224 mm and average length
0.491 mm. Frontal wall formed by 15 to 18 costae and lateral
costal fusions producing 8 to 10 pores between each two costae.
Median area small, porous. Aperture semilunar, narrow. No oral
spines Avicularia large, intrazooecial, with long rostrate and
pivotal bar. Hyperstomial ovicell semilunar, narrow, strongly
convex, smooth, nonporous with smooth, low, median rib.

The described specimens are very similar in shape

of the ovicells and construction of the frontal shield to the
holotype deposited in the Natural History Museum in Vien-
na. The size of the holotype is little larger (0.32


0.48 mm).

However Schmid (1989) shows range of width of the zooeci-
um from 0.20 to 0.48 mm and range of length of zooecium
from 0.32 to 0.52 mm.

Štrba locality, with one probable specimen

from Hybica locality.

Family Pelmatoporidae Lang 1916

Subfamily Castanoporinae Lang 1916

Genus Anornithopora Lang 1916

Anornithopora pretiosa 

(Reuss 1866)

(Pl. II: Fig. 2)

1866 Lepralia pretiosa sp.n., Reuss p. 175, Pl. 8, Fig. 4
1994 Anornithoporapolygona Voigt, Zágoršek p. 372, Fig. 6c


Zoaria encrusting. Zooecia cribrimorph, oval,

small. Frontal wall slightly convex, with about 20 costae. Ar-
eolar pores small, clearly visible. The costae form wrinkles
in the median area of frontal wall. The median area is more
than half as long again as the length of zooecium. Gymno-
cyst usually missing. Orifice oval, with wide peristome. No
secondary aperture. Apertural bars wide, straight. No avicu-
laria. Ovicells have not been observed.

The described specimens are identical with Lep-

ralia pretiosa 

described by Reuss 1866. According to Lang

(1922) the genus Anornithopora has small zooecia, with

Plate II:

 Fig. 1. Cribrilaria haueri (Reuss 1848) — Štrba locality,

SEM BS 300, photo by I. Holický. Fig. 2. Anornithopora pretiosa
(Reuss 1866) — Hybica locality, SEM BS 300, photo by I. Holický.
Figs. 3–4. Scrupocellaria brendolensis 

Waters 1891 — Štrba locali-

ty, SEM BS 300, photo by I. Holický. (Fig. 4 detail of the frontal
avicularium.) Fig. 5. Lunulites quadrata (Reuss 1848) — Východná
locality, photo by the author. Figs. 6–7. Cribrilaria radiata (Moll
1803) — Hybica locality, SEM BS 300, photo by I. Holický. Scale
bars represent 1 mm (Fig. 1 and 5) and 0.1 mm (remaining figures).

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408                                                                                              ZÁGORŠEK


Braga Gp., 1963: I Briozoi del Terziario Veneto. Boll. Soc. Pal. It.,

2, 1, 16–55.

Braga Gp., 1975: Briozoi dell’Eocene di Possagno. Schweiz.

Paläont. Abh., 

97, 141–148.

Braga Gp., 1980: Paleontology. In: Antolini P., Braga G.& Finotti

F. (Eds.): I Briozoi dei dintorni di Rovereto. Monte Baldo set-
tentrionale e Valle di Gresta. Mus. Civ. Rovereto, 

82, 1–103.

Braga Gp., 1991: Reuss‘ collection of Cheilostome Bryozoans

from Venetia stored in Natur-Historisches Museum Wien: A
Proposed Revision. In: Bigey: Bryozoa living and fossil. Bull.
Soc. Sci. Nat. Ouest Fr., Mem. HS, 

1, 49–59.

Braga Gp. & Barbin G., 1988: Les Bryozoaries du Priabonien

Stratotypique (Province Vicenza, Italie). Revue de Paleobiol-

7, 2, 495–556.

Braga Gp. & Munari M., 1972: Studi biometrici su due popolazio-

ni di Conescharellina (C. perfecta Accordi e C. veronensis
Accordi) dell’Eocene superiore del Veneto. Studi Trent. Sc.
Nat. sec. A, 

49, 111–126.

Canu F. & Bassler R.S., 1920: North American Early Tertiary Bry-

ozoa. U.SNat. Mus. Bull., 106, 1–879.

David L. & Pouyet S., 1974: Revision des Bryozoaires Cheilos-

tomes miocenes du Bassin de Vienne (Autriche). Doc. Lab.
Geol. Fac. Si. Lyon, 

60, 83–257.

Gordon D., 1984: The Marine Fauna of New Zealand: Bryozoa:

Gymnolaemata from the Kermadec Ridge. New Zealand
Oceanographic Institute Memoir 91, Wellington, 


Gordon D., 1986: The Marine Fauna of New Zealand: Bryozoa: Gym-

nolaemata (Ctenostomata and Cheilostomata Anasca) from the
Western South Island Continental Shelf and Slope. New Zealand
Oceanographic Institute Memoir 95, Wellington,


Gross P., Köhler E. & Samuel O., 1984: New lithostratigraphical

division of Inner-Carpathian Paleogene. GeolPráce, Spr.,
81, 103–117 (in Slovak).

Gross P. & Köhler E., Eds., 1980: Geology of Liptov Basin.


 Bratislava, 1–217 (in Slovak).

Lang 1922: Catalogue of the Fossil Bryozoa (Polyzoa). Vol. IV

The Cretaceous Bryozoa (Polyzoa), the Cribrimorphs part II.
British Museum (Natural History),

 London, 1–404 .

Reuss A.E., 1848: Die fossilen Polyparien des wiener Tertiärbeck-

Table 1:

 List of the Anascan Bryozoa from the Western Carpathains (Slovakia).

T a x a







L a s t   d e s c r i p t i o n

A ld e r in a   s u b ti lim a r g o   ( R e u s s   1 8 6 4 )






Z á g o r š e k   1 9 9 6 a ,  p .  1 2 3 ,  P l .  I ,  F i g .  3   -   5

A n o r n it h o p o r a     p r e t i o s a   ( R e u s s   1 8 6 6 )



B iflu s t r a    s a v a r t i i    t e x t u r a t a   ( R e u s s   1 8 4 8 )



Z á g o r š e k   1 9 9 6 a ,  p .  1 2 3 ,  P l .  I ,  F i g .  1   -   2

C a lp e n s ia     g r a c i l i s   (M ü n ster  in   G o ld fu ss  1 8 2 6 )



Z á g o r š e k   1 9 9 6 b ,  p .  5 2 9 ,  P l .  I ,  F i g .  7

C a lp e n s ia     h e x a g o n a   Z á g o r š e k   1 9 9 4



Z á g o r š e k   1 9 9 6 b ,  p .  5 3 1 ,  P l .  I V ,  F i g .  1 ,  2 ,  4

C a lp e n s ia     p o ly s t i c h a   ( R e u s s   1 8 4 8 )


* ?


Z á g o r š e k   1 9 9 6 b ,  p .  5 3 1 ,  P l .  I I I ,  F i g .  7

C e ll a r ia     r e u s s i     d ' O r b i g n y   1 8 5 1




C r a s s im a r g in a t e l l a     m a c r o s t o m a  (R euss 1848)







Z á g o r š e k   1 9 9 6 a ,  p .  1 2 3 ,  P l .  I ,  F i g .  6 ,  P l .  I I ,  F i g .  1

C r i b r ila r i a     h a u e r i   ( R e u s s   1 8 4 8 )




C r i b r ila r i a     r a d i a t a   ( M o l l   1 8 0 3 )



C h a p e r ia     s p in e l l a     Z á g o r š e k   1 9 9 4



Z á g o r š e k   1 9 9 6 a ,  p .  1 2 5 ,  P l .  I I ,  F i g .  4   -   7

C h lid o n io p s is     v i n d o b o n e n s i s   ( R e u s s   1 8 4 8 )



L u n u li te s    q u a d r a t a   ( R e u s s   1 8 4 8 )



O n y c h o c e ll a     s u b p y r i f o r m i s   (d 'A rc h ia c  1 8 4 6 )







Z á g o r š e k   1 9 9 6 b ,  p .  5 2 5 ,  P l .  I ,  F i g .  1   -   6

P u e l lin a     ( C r ib r i l a r i a )     s c r i p t a   (R e u ss 1 8 4 8 )

* ?



A m p h ib le s tr u m     a p p e n d ic u l a t a   (R e u s s   1 8 4 8 )

* ?

* ?



R o s s e lia n a     r o s s e l i i   ( A u d o u i n   1 8 2 6 )


Z á g o r š e k   1 9 9 6 b ,  p .  5 2 9 ,  P l .  I I I ,  F i g .  6

S c r u p o c e ll a r ia     b r e n d o l e n s i s     W a t e r s   1 8 9 1



S m ittip o r a     g r a n d i c o n i s   Z á g o r š e k   1 9 9 6


Z á g o rše k   1 9 9 6 b ,  p .  5 2 7 ,  P l.  II,  F ig .  1   -  5 ,  P l.  IV ,  F ig .  3

S m ittip o r a     te n u i s   ( C a n u   &   B a s s l e r   1 9 2 0 )


Z á g o r š e k   1 9 9 6 b ,  p .  5 2 7 ,  P l .  I I I ,  F i g .  1   -   3

S t e g i n o p o r e lla     c e l l a r i i f o r m i s   C h eeth am   1 9 6 3


Z á g o r š e k   1 9 9 6 b ,  p .  5 3 3 ,  P l .  I V ,  F i g .  5

S t e g i n o p o r e lla     h a i d i n g e r i   ( R e u s s   1 8 4 8 )





Z á g o r š e k   1 9 9 6 b ,  p .  5 3 1 ,  P l .  I V ,  F i g .  6   -   7

V ib r a c e lla     tr a p e z o i d e a   ( R e u s s   1 8 4 8 )



Z á g o r š e k   1 9 9 6 b ,  p .  5 2 9 ,  P l .  I I I ,  F i g .  4

few costae (20 or fewer), no secondary aperture and very
few or no avicularia. The described specimens are coherent
with the genus Anornithopora in forming a median lamella,
the apertural bar and in the absence of avicularia.

The new specimens from Štrba allow a more precise deter-

mination than the specimens from Hybica (Zágoršek 1994).

Štrba and Hybica localities


Six Eocene localities with Bryozoa have been newly dis-

covered in the Slovak part of the Western Carpathians. A to-
tal of 24 Eocene anascan Bryozoa (see Table 1) are known
from the Slovak part of the Western Carpathians. The richest
bryozoan associations are found at the Hybica and Štrba lo-
calities which have similar lithology. They also have the
most species in common (six).

The environment of the localities is of two different types.

First, shallow marine with Lunulites (and Lacrimula) where
Bryozoa are common (Východná, Partizánska  upča localities,
and probably also Rajecké Teplice and Poluvsie localities).

The second, neritic, margin of the shelf influenced by up-

welling, where the bryozoan zoaria are only represented by
allochems in the sediments (Zágoršek 1996).


The field work was funded by AAPG

Grant (Grant-in-Aid 582-12-01). Cost of SEM photography
and the final version of the plates was reimbursed by
VEGA grant number 4080. Great thanks also go to Prof.
Vávra from the Department of Paleontology Vienna Univer-
sity who allowed me to study the original material of Reuss
and Stoliczka species in Vienna Natural History Museum
and helped me with his comments. I would also like to ex-
press my sincere thanks to Prof. Braga who helped me very
much with their kind comments and improved the quality of
this paper.

background image


ens. Ein monographischer Versuch. Naturwiss. Abhadlung., 2,
1, 1–109.

Reuss A.E., 1866: Die Foraminiferen, Anthozoen und Bryozoen

des deutschen Septarienthones. Ein Beitrag zur Fauna der
mitteloligocänen Tertiärschichten. Denkschr. k. Akad. Wiss.,
Math.-Naturwiss. Cl., 

25, 117–214.

Reuss A.E., 1869: Paläontologische Studien über die älteren Ter-

tiärschichten der Alpen. 2) Die fossilen Anthozoen und Bryo-
zoen der Schichtengruppe von Crosara. Denkschr. k. Akad.
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29, 215–294.

Schmid 1989: Cheilostome Bryozoen aus dem Badenien (Miozän)

von Nußdorf (Wien). Beiträge zur Paläontologie von Österre-

15, 1–70.

Waters A.W., 1891: North-Italian Bryozoa. Quart. JlGeol. Soc.,

47, 1–34.

Zágoršek K., 1994: Late Eocene (Priabonian) Cheilostomata Bryo-

zoa from Liptov Basin — Western Carpathians (Slovakia). N.

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, 413–422.

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Western Carpathians. Miner. slovaca, 28, 120–128 (in Slovak).

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ern Carpathians. Miner. slovaca, 28, 523–534 (in Slovak).