GEOLOGICA CARPATHICA, 48, 1, BRATISLAVA, FEBRUARY 1997
49–61
JURASSIC RADIOLARITES FROM THE EASTERN PART
OF THE PIENINY KLIPPEN BELT
(WESTERN CARPATHIANS)
LADISLAVA OŽVOLDOVÁ and LENKA FRANTOVÁ
Department of Geology and Paleontology, Faculty of Sciences, Comenius University, Mlynská dolina,
842 15 Bratislava, Slovak Republic
(Manuscript received May 9, 1996; accepted in revised form December 12, 1996)
Abstract:
The radiolarites of the Šariš and Pieniny part of the Pieniny Klippen Belt belong to the Pieniny Succesion
s.l. (Nemčok et al. l990). A horizon of radiolarites from the locality Milpoš and Podsadek near Stará ubovňa contains
a considerable calcareous admixture. At the localities of Šarišské Jastrabie and Lúčka the radiolarites are almost without
calcite and contain Mn coatings. The radiolarian associations represent UAZone 8 – UAZone l0 ranging from middle
Callovian to early Kimmeridgian (sensu Baumgartner et al. l995). Radiolarites with Mn coatings reach to middle
Oxfordian at the locality Šarišské Jastrabie. A new genus Fultacapsa nov. gen. is described in this paper.
Key words:
Jurassic, Western Carpathians, Pieniny Klippen Belt, Pieniny Succession, radiolarians, radiolarites.
Introduction
Up to now the occurrence of radiolarites in the eastern part of
the Pieniny Klippen Belt has not been proved by evidence of
radiolarian microfauna. In this first contribution we deal with
the evaluation of radiolarian microfauna at the localities of
Milpoš, Lúčky, Kyjov, Šarišské Jastrabie (the Šariš part of
the Klippen Belt) and Podsadek near Stará ubovňa (the Pi-
eniny part of the Klippen Belt) (Fig. 1).
On the Polish territory Birkenmajer (l977) determined the
lithostratigraphical units for the Pieniny Klippen Belt.
For radiolarites they are as follows:
l — Sokolica Radiolarite Formation — grey-green, dark-grey
or black radiolarites with the typical coatings of Mn minerals.
The age — ?Upper Bajocian-Callovian – ?Lower Oxfordian.
2 — Czajakowa Radiolarite Formation with three members:
lower red radiolarites — Kamionka Radiolarite Member
green radiolarites — Podmajerz Radiolarite Member
upper red radiolarites — Buwald Radiolarite Member
Age — ?Upper Callovian-Upper Oxfordian-?Kimmeridgian.
The latest comprehensive geological research in the inves-
tigated area was carried out by Nemčok (l982, l986) and
Nemčok et al. (l990).
In accordance with this information radiolarites belong to
the Pieniny Succession s.l. They overlie the crinoidal or spot-
ty limestones. In the underlying strata, nodular limestones or
the pelagic facies occur.
Within the research of Nemčok et al. (l986) the investiga-
tion of radiolarian microfauna was also performed for the lo-
calities of Podsadek and Šarišské Jastrabie by Ondrejíčková
(l985). The age for radiolarian associations from radiolarites
was determined as the Upper Jurassic.
Fig. 1.
Location of the investigated area and sampling localities.
a
— Situation of investigated area (A). b — Close view of the in-
vestigated area (A). c — Podsadek section — Ps. d — Šarišské
Jastrabie sections — ŠJP, ŠJ, ŠJDD. e — Kyjov area — K. f —
Lúčka section — L, Milpoš section — M.
50 OŽVOLDOVÁ and FRANTOVÁ
Fig. 2.
Lithological columns of the sections ŠJP, ŠJDD and ŠJ (lo-
cality Šarišské Jastrabie).
Stratigraphy of investigated localities
Milpoš (Fig. 1f)
Radiolarites overlie the red crinoidal limestones some-
times with a layer or lense of cherts (Krupianka Limestone
Formation sensu Birkenmajer l977). Except for crinoidal
plates they contain fragments of forams, ostracods and phan-
toms of radiolarians. There is a debris section between the
crinoidal limestones and the overlying radiolarites. The sec-
tion continues with the rusty-red radiolarites of the Czaja-
kowa Radiolarite Formation (Buwald Radiolarite Member),
sometimes with the occurrence of the green parts.
The radiolarian microfauna from radiolarites was very
poor. Only one sample (M-5) has been analyzed, in which
the species Fultacapsa sphaerica (Ožvoldová) nov. comb.
occurred. Its presence shows that the association is not older
than UAZone 9 ranging from middle to late Oxfordian (sensu
Baumgartner et al. l995, as well as the results of our investi-
gations).
In the upper part of the radiolarite horizon a great number
of intercalations of pinkgrey limestones appear. They become
dominant into the overlying strata and they include numerous ir-
regular intercalations of siliceous nodules. The limestones only
contain phantoms of radiolarians, filled up with chalcedony or
calcite. The phantoms of radiolarians are less numerous in the
siliceous intercalations. The presumptive age of this part of the
bed is Kimmeridgian. The Tithonian pelagic limestones (Pien-
iny Limestone Formation sensu Birkenmajer l977) with rich mi-
crofauna overlie these limestones.
Lúčka (Fig. 1f)
In the lower part of the outcrop of radiolarites, green-grey
to dark-grey radiolarite layers with Mn coatings occur
(Sokolica Radiolarite Formation). Grey-green and rusty-red
radiolarites overlie them (Podmajerz Radiolarite Member
and Buwald Radiolarite Member of Czajakowa Radiolarite
Formation).
The poor radiolarite association from the rusty-red radi-
olarite (sample L-5) represented a middle Oxfordian – early
Kimmerdgian age (the species Fultacapsa sphaerica
(Ožvoldová) nov. comb. and Zhamoidellum ovum Dumitrica
appear in UAZone 9 ranging from middle to late Oxfordian
and the species Emiluvia premyogii Baumgartner finishes in
UAZone 10 — Oxfordian–early Kimmeridgian according to
Baumgartner et al. l995).
Kyjov (Fig. 1e)
The grey-green and rusty-red radiolarites occur in the scree
material only.
The grey-green radiolarite, sample K-2, contained a rela-
tively poor association. The presence of the species
Podobursa spinosa
(Ožvoldová) proves that the assemblage
is not older than UAZone 8 ranging from middle Callovian to
early Oxfordian (l.c.).
In the rusty-red radiolarite the radiolarian association, pre-
sented in sample K-1 corresponds to UAZone 8–l0 — mid-
dle Callovian–early Kimmeridgian (l.c.). The species
Podobursa spinosa
(Ožvoldová) and Emiluvia orea orea
Baumgartner, which appear in UAZone 8 and Paronaella
mulleri
Pesagno and Angulobracchia digitata Baumgartner,
which finish in UAZone l0 occur in this association.
Concerning the stratigraphical range of Emiluvia orea orea
Baumgartner, this species does not occur with such species as
Guexella nudata
(Kocher) and Stylocapsa oblongula Kocher
(UAZone 8) according to our investigations. It appears later.
Šarišské Jastrabie (Fig. 1d, Fig. 2)
This locality is situated in the eastern part of the village in
the brook Vesné. Three sections have been ivestigated in this
area: ŠJ, ŠJP and ŠJDD.
The section ŠJ consists of outcrops of radiolarites of grey-
green to dark-grey colour, partly of reddish colour with shale
intercalations of similar colour. The radiolarite levels some-
times show coatings of Mn minerals (Sokolica Radiolarite
Formation). The radiolarian microfauna was very poor, most-
ly indeterminable.
The section ŠJP presents brown-grey and green-grey radi-
olarites in its lower part, sometimes with rusty-red parts and
with irregular pigmentation of Mn minerals (Sokolica Radi-
olarite Formation). The radiolarite levels are separated by
thin shale intercalations.
The sample ŠJP-1, in the lowermost part of the section
yielded a badly preserved association with only one deter-
minable species Eucyrtidiellum ptyctum Riedel & Sanfilip-
po. According to Baumgartner et al. (l995) this species ap-
pears in UAZone 5, ranging from latest Bajocian to early
Bathonian. We agree with the opinions presented by Baum-
gartner (l984) and Goričan (l994) that this species appears
during the Callovian.
JURASSIC RADIOLARITES FROM EASTERN PART OF PIENINY KLIPPEN BELT 51
Approximately 0.5 m upwards a richer association occurs
in sample ŠJP-2a, which contains the species Emiluvia pes-
sagnoi multipora
Steiger. This species appears in UAZone 8,
ranging from middle Callovian to early Oxfordian (Baum-
gartner et al. l995) and co-occurs with the species Willir-
iedellum
sp. A sensu Matsuoka l983, which makes its last
occurrence in this zone. These data indicate that the associa-
tion represents a middle Callovian–early Oxfordian age.
In the upper part of the outcrop the Mn coatings disappear.
Green-grey or grey-green radiolarites with reddish parts oc-
cur there (Podmajerz Radiolarite Member of Czajakowa Ra-
diolarite Formation).
The uppermost part of the outcrop is formed by rusty-red
radiolarites (Buwald Radiolarite Member of the Czajakowa
Radiolarite Formation). The rich association found in sample
ŠJP-4 contained two types of radiolarian tests: a) well pre-
served, often pellucid (fluoritization) and b) worse preserved
with Fe pigmentation, often with features of pressure defor-
mation. The second type seems originate in argillaceous lam-
inae, which are probably formed by bottom currents (the
signs of the shape sorting). The radiolarian tests in the
argillaceous parts are more sensitive to deformation during
the compaction of sediment.
In the sample ŠJP-4 the following species occur: Podocapsa
amphitreptera
Foreman, Fultacapsa sphaerica (Ožvoldová),
nov. comb., ranging from middle to late Oxfordian (UAZone 9
of Baumgartner et al. l995) and the species Tritrabs casmali-
aensis
(Pessagno), which has its last occurrence in UAZone
10, ranging from late Oxfordian to early Kimmeridgian (l.c.).
The above mentioned data indicate that the association repre-
sents a middle Oxfordian–early Kimmeridgian age.
In this section chemical analyses (Fig. 3) were performed on
the grey radiolarites (ŠJP-l), the grey-green radiolarites with Mn
coatings (ŠJP-2), the green superposed radiolarites (ŠJP-3) and
the rusty-red radiolarites at the top of the section (ŠJP-4).
The composition of CaO increases towards the top of the
section but if we compare with the radiolarites of the western
part of the Klippen Belt, these values are very low ( max.
0.36 %). The concentration of Al
2
O
3
and Fe
2
O
3
increases up-
wards. In the radiolarites with Mn coatings (ŠJP-2) the con-
centration of Mn (3.l6 %) highly exceeds the values obtained
from the underlying and overlying beds (ŠJP-1 — l38 ppm,
ŠJP-3 — 28l ppm, ŠJP-4 — 326 ppm).
According to the research of Ilavský (1955) the enriched
radiolarites with Mn contained as much as 18 % Mn.
On the right side of the outcrop a tectonic contact between the
Sokolica Radiolarite Formation and the Eocene Globigerinian
marls occurs. These marls are of dark-grey to black colour with
rich pigmentation of Mn minerals. Their presumptive age was
Middle Jurassic (Ilavský 1955), but later investigations proved
that they are of Eocene age (Gašparíková l986).
In the other places of this locality the radiolarites are over-
lain by red nodular limestones of Kimmeridgian-Early Titho-
nian age (Doležalová l993). This fact demonstrates the corre-
spondence of this section to the Kysuca Succession.
In the section ŠJDD, overturned beds of radiolarites with
thin intercalations of shales occur. In the middle part of the
outcrop, a tectonic contact between radiolarites and dark-
grey and black shales containing rich impregnations of Mn
pigment and intraclasts of light soft clayey material with
high concentration of P
2
O
5
(8.29 %) occurs. Paleontological
evidence for their stratigraphical assignment does not exist.
In the top part of the outcrop the radiolarian fauna from
dark-grey to green-grey radiolarites with Mn coatings
(Sokolica Radiolarite Formation) was very poor. The pres-
ence of the species Podocapsa amphitreptera Foreman (sam-
ple ŠJDD-4) demonstrates that the association is not older
than UAZone 9, ranging from middle to late Oxfordian
(Baumgartner et al. l995).
The underlying grey-green to green radiolarites, sample
ŠJDD-5, (Podmajerz Radiolarite Member of Czajakowa Ra-
diolarite Formation) contained the species Fultacapsa
sphaerica
(Ožvoldová) nov. comb., which appears in UA-
Zone 9, ranging from middle to late Oxfordian (l.c.). In the
lower part, in rusty-red levels, the species Emiluvia orea
Baumgartner in sample ŠJDD-8 occurred. In the bottom of
the outcrop the rusty-red radiolarites contained an associa-
tion (sample ŠJDD-9) which represents UAZone 9–UAZone
10, ranging from middle Oxfordian to early Kimmeridgian
(l.c.) according to the co-occurrence of the species Fultacap-
sa sphaerica
(Ožvoldová) nov. comb., (first occurrence in
UAZone 9) and Tritrabs casmaliaensis (Pessagno) (late oc-
currence in UAZone l0).
The above mentioned data resulted in the conclusion that
the radiolarites in this outcrop represent a middle Oxfordian-
early Kimmeridgian age.
Podsadek near Stará ubovňa (Fig. 1c)
The section is situated in the brown-grey siliceous limestones
with grey cherts in the lower part and the rusty-red or green sili-
ceous limestones to calcareous radiolarites in the upper part of
the outcrop. The Middle-Upper Tithonian Rogoznik limestones
overlie these strata (Nemčok et al. 1982).
In the brown-grey limestones the chert contained an assem-
blage (sample Ps-3) representing middle Oxfordian–early
Fig. 3.
Chemical analyses of the samples from the section ŠJP (lo-
cality Šarišské Jastrabie).
1
P
J
P
J 2
P
J 3
P
J 4
H
2
O
+
)
%
(
9
0
.
0
8
0
.
0
8
0
.
0
9
0
.
0
H
2
O
)
%
(
7
9
.
1
8
7
.
5
1
8
.
4
9
5
.
3
O
i
S
2
)
%
(
9
4
.
4
9
5
9
.
4
8
7
9
.
0
8
6
9
.
1
8
O
i
T
2
)
%
(
0
1
.
0
9
0
.
0
1
1
.
0
1
1
.
0
l
A
2
O
3
)
%
(
1
4
.
1
1
2
.
2
0
8
.
6
9
5
.
6
e
F
2
O
3
)
%
(
4
9
.
0
3
7
.
1
6
8
.
3
5
3
.
4
)
%
(
O
a
C
9
1
.
0
8
2
.
0
6
2
.
0
6
3
.
0
)
%
(
O
g
M
8
1
.
0
0
3
.
0
9
4
.
1
2
2
.
1
K
2
)
%
(
O
5
2
.
0
8
2
.
0
6
2
.
1
8
3
.
1
a
N
2
)
g
k
/
g
m
(
O
0
6
.
3
4
9
0
7
.
5
3
9
0
9
.
4
0
2
1
0
1
.
3
0
2
1
)
g
k
/
g
m
(
n
M
3
8
.
8
3
1
0
0
.
0
0
6
1
3
5
2
.
1
8
2
2
2
.
6
2
3
)
g
k
/
g
m
(
r
C
7
0
.
1
1
5
0
.
1
1
7
9
.
7
3
2
9
.
2
3
)
g
k
/
g
m
(
i
N
6
0
.
0
1
7
0
.
6
1
7
4
.
6
3
2
9
.
0
3
)
g
k
/
g
m
(
u
C
4
2
.
0
4
2
2
.
9
4
9
8
.
6
1
1
9
8
.
3
4
52 OŽVOLDOVÁ and FRANTOVÁ
Kimmeridgian (UAZone 9–10) (Baumgartner et al. l995) on
the basis of the co-occurrence of Emiluvia premyogii Baum-
gartner and Orbiculiforma (?) catenaria Ožvoldová with
Emiluvia ordinaria
Ožvoldová.
In the upper part the radiolarian assemblage from the
rusty-red calcareous radiolarite (sample Ps-14) contained the
species Paronaella pristidentata Baumgartner, which has its
first occurrence in UAZone 10 (late Oxfordian to early Kim-
meridgian) and the species Paronaella mulleri Pessagno,
which finishes in this zone (l.c.). The association represents a
late Oxfordian–early Kimmeridgian age (UAZone 10 of
Baumgartner et al. l995).
The horizon of radiolarites in this locality has a great quan-
tity of calcareous component. According to this research the
most siliceous part of the section represents the middle Ox-
fordian–early Kimmeridgian interval.
Summary of the results obtained
Radiolarites from the studied localities belong to the Pien-
iny Succession s.l. (Nemčok et al. 1990). In the localities of
Milpoš and Podsadek near Stará ubovňa the horizon of ra-
diolarites includes a large quantity of limestone layers and
the radiolarites have mostly fair calcareous admixture. The
strata with regard to the underlying crinoidal limestones be-
long to the Czertezik Succession.
In the locality of Šarišské Jastrabie the nodular limestones
overlie radiolarites (Doležalová l993). This fact demon-
strates correspondence to the Kysuca Succession.
The underlying and overlying rocks to the radiolarite hori-
zon are not uncovered in the locality of Lúčka. However, the
radiolarites are similar to those in the locality of Šarišské
Jastrabie.
The radiolarites in the locality of Kyjov only occur in the scree.
The important feature of the Sokolica Radiolarite Forma-
tion in the localities of Šarišské Jastrabie and Lúčka is the
presence of Mn coatings. Ilavský (l955) gives information
concerning the concentrations of Mn minerals in radiolarites
in the Vesné brook of the Šarišské Jastrabie (section ŠJP).
This author found that the course and the amount of these
concentrations was irregular. The richest zone is located near the
contact with the shales, which are also covered by manganesef-
erous coating and are Middle Jurassic in age, according to Ilavský
(l955). He assumed the migration of Mn minerals towards the
contact with the impermeable horizon of shales.
Recent research proved that the contact with these shales is
tectonic and the shales belong to the Paleogene series (Proč
beds) of the Klippen Belt (Nemčok et al. 1990).
The presence of Mn in radiolarites through the migration
from the original source can be supported by the preliminary
chemical analyses presented in this paper and carried out in the
section ŠJP in Šarišské Jastrabie. While the concentration of Mn
in the dark-grey radioarite with Mn coatings in the lower part of
the section (sample ŠJP-2) was 3.16 %, in the underlying grey
beds (sample ŠJP-1) and overlying green and rusty-red radiolar-
ites (samples ŠJP-3 and ŠJP-4) the concentration of Mn was low
(138 ppm, 281 ppm, 326 ppm).
It can be supposed that the accumulation of Mn minerals
was realized in the course of the migration from the original
source into the radiolarite beds originally having plenty of
joints. Under the microscope the original siliceous matter is
sometimes broken into angular parts separated by chlorite-
siliceous fissures with Mn pigment.
Microscopic analysis of the black shales from the section
ŠJDD show that the rock is thickly impregnated with small glob-
ular particles of Mn minerals covering thickly the original rock.
The analyses of microfauna show well preserved radiolari-
ans in Šarišské Jastrabie. In the lower part of the radiolarite
horizon the radiolarian assemblages represent the middle
Callovian–early Oxfordian interval (UAZone 8 sensu Baum-
gartner et al. l995). The upper part of the radiolarite horizon
is correlatable to the UAZone 9–UAZone 10 (middle Oxfor-
dian–early Kimmeridgian) (l.c.).
Plate I: Fig. 1.
Archaeospongoprunum imlayi Pessagno — ŠJP-2a,
8151, 150
×
. Fig. 2. Transhsuum brevicostatum (Ožvoldová) —
ŠJP-2a, 1860, 240
×
. Fig. 3. Tritrabs rhododactylus Baumgartner
— ŠJP-2a, 1906, 170
×
. Fig. 4. Emiluvia pessagnoi multipora
Steiger — ŠJP-2a, 8129, 165
×
. Fig. 5. Triactoma blakei (Pessag-
no) — ŠJP-2a, 1921, 160
×
. Fig. 6. Bernoullius dicera (Baumgart-
ner) — ŠJP-2a, 1867, 180
×
. Fig. 7. Paronaella kotura Baumgart-
ner — ŠJP-2a, 8147, 130
×
. Fig. 8. Orbiculiforma (?) cf. catenaria
Ožvoldová — ŠJP-2a, 8120, 160
×
. Fig. 9. Paronaella broenni-
manni
Pessagno — ŠJP-2a, 2383, 140
×
. Fig. 10. Homoeopa-
ronaella argolidensis
Baumgartner — ŠJP-2a, 8133, 150
×.
Fig. 11.
Paronaella mulleri Pessagno — ŠJP-2a, 8137, 175
×
.
Fig. 12.
Triactoma sp. — ŠJP-2a, 8106, 240
×
. Fig. 13. Triactoma
jonesi
(Pessagno) — ŠJP-2a, 8156, 160
×
. Fig. 14. Sethocapsa sp.
— ŠJP-2a, 1882, 290
×
.
Plate II: Fig. 1.
?Dictyomitrella sp. — ŠJP-2a, 8145, 360
×
. Fig. 2.
Parvicingula dhimenaensis
Baumgartner — ŠJP-2a, 2387, 250
×
.
Fig. 3
. Archaeodictyomitra directiporata (Rüst) — ŠJP-2a, 1917,
280
×
. Fig. 4. Cinguloturris carpatica Dumitrica — ŠJP-2a, 1883,
260
×
. Fig. 5. Williriedellum sp. A sensu Matsuoka,1983 — ŠJP-2a,
1875, 360
×
. Fig. 6. Eucyrtidiellum ptyctum (Riedel & Sanfilippo) —
ŠJP-2a, 8108 490
×
. Fig. 7. Archaeodictyomitra prisca Kozur & Mos-
tler — ŠJP-2a, 8107, 280
×
. Fig. 8. Pseudodictyomitrella hexagonata
(Heitzer) — ŠJP-2a, 8095, 450
×
. Fig. 9. Transhsuum maxwelli (Pes-
sagno) — ŠJP-2a, 1897, 230
×
. Fig. 10. Mirifusus cf. fragilis Baum-
gartner — ŠJP-2a, 8151, 150
×
. Fig. 11. Tricolocapsa yaoi Matsuoka —
ŠJP-2a, 8102, 400
×
. Fig. 12. Tricolocapsa undulata (Heitzer) — ŠJP-2a,
1894, 410
×
.
Plate III: Fig. 1.
Acanthocircus suboblongus (Yao) — ŠJP-4, 8330,
210
×
. Fig. 2. Orbiculiforma (?) catenaria Ožvoldová — ŠJP-4, 8306,
160
×
. Fig. 3. Archaeospongoprunum imlayi Pessagno — ŠJ DD-9,
4515, 220
×
. Fig. 4. Archaeospongoprunum sp. — K-1, 4570, 165
×
Fig. 5.
Emiluvia orea Baumgartner, the side view of Fig. 6 — Ps-3,
7010, 170
×
. Fig. 6. Emiluvia orea Baumgartner — Ps-3, 7009, 145
×
.
Fig. 7.
Paronaella pygmaea Baumgartner — ŠJ DD-9, 4514, 145
×
.
Fig. 8.
Emiluvia ordinaria Ožvoldová — Ps-3, 5831, 150
×
. Fig. 9.
Hexastylus
sp. — Ps-3, 7001, 130
×
. Fig. 10. Triactoma blakei (Pessa-
gno) — Ps-3, 5836, 140
×
. Fig. 11. Paronaella broennimanni Pessag-
no — K-1, 4576, 190
×
. Fig. 12. Fultacapsa cf. sphaerica (Ožvoldová)
nov. comb. — PS-l4, 2341, l80
×
. Fig. 13. Archaeospongoprunum im-
layi
Pessagno — Ps-14, 4265, 225
×
. Fig. 14. Emiluvia premyogii
Baumgartner — Ps-3, 5838, 175
×
. Fig. 15. Paronaella mulleri Pessa-
gno — K-1, 4581, 135
×
. Fig. 16. Triactoma jonesi (Pessagno) —
ŠJP-4, 8331, 190
×
. Fig. 17. Halesium sp. — K-1, 4568, 120
×
.
Fig. 18.
Angulobracchia cf. rugosa Jud — Ps-14, 3060, 145
×
.
Fig. 19.
Emiluvia sedecimporata
(Rüst) — Ps-14, 3048, 145
×
.
§
PLATE I 53
54 PLATE II
PLATE III 55
56 PLATE IV
PLATE V 57
58 PLATE VI
JURASSIC RADIOLARITES FROM EASTERN PART OF PIENINY KLIPPEN BELT 59
Plate IV: Fig. 1.
Orbiculiforma (?) catenaria Ožvoldová — Ps-3,
2852, 125
×
. Fig. 2. Tritrabs rhododactylus Baumgartner — K-2,
4507, 120
×
. Fig. 3. Tetraditryma pseudoplena Baumgartner — K-1,
4579, 110
×
. Fig. 4. Archaeodictyomitra prisca Kozur & Mostler —
ŠJ DD-6, 4499, 250
×
. Fig. 5. Transhsuum brevicostatum
(Ožvoldová) — K-3, 4563, 250
×
. Fig. 6. Emiluvia chica Foreman —
Ps-3, 5845, 160
×
. Fig. 7. Haliodictya (?) hojnosi Riedel & Sanfilip-
po — Ps-14, 4277, 220
×
. Fig. 8. Syringocapsa spinellifera Baumgart-
ner — ŠJP-4, 8352, 170
×
. Fig. 9. Haliodictya (?) antiqua
(Rüst) — ŠJP-4, 8327, 95
×
. Fig. 10. Emiluvia salensis Pessagno —
Ps-14, 3028, 140
×
. Fig. 11. Emiluvia pessagnoi Foreman — Ps-14,
4287, 130
×
. Fig. 12. Tritrabs exotica (Pessagno) — ŠJP-4, 1574,
195
×
. Fig. 13. Tetratrabs bulbosa Baumgartner — Ps-14, 3014, 60
×
.
Fig. 14.
Perispyridium ordinarium Pessagno — Ps-14, 4275, 170
×
.
Fig. 15.
Zhamoidellum ovum Dumitrica — ŠJ DD-5, 4492, 420
×
.
Fig. 16.
Parvicingula cf. dhimenaensis Baumgartner — ŠJP-4, 8304,
240
×
. Fig. 17. Parvicingula cf. mashitaensis Mizutani — Ps-14,
4282, 300
×
. Fig. 18. Pseudodictyomitra cf. primitiva Matsuoka &
Yao — Ps-14, 4273, 210
×
.
Plate V: Fig. 1.
Fultacapsa sphaerica (Ožvoldová) nov. comb. —
ŠJP-4, 1579, 210
×
. Fig. 2. Fultacapsa sphaerica (Ožvoldová) nov.
comb. — Ps-14, 3039, 140
×
. Fig. 3. Fultacapsa aff. sphaerica
(Ožvoldová) nov. comb. — Ps-14, 3051, 195
×
. Fig. 4. Podobursa
polyacantha
(Fischli) — ŠJP-4, 8346, 170
×
. Fig. 5. Podobursa spi-
nosa
(Ožvoldová) — Ps-3, 5837, 130
×
. Fig. 6. Podocapsa amphi-
treptera
Foreman — Ps-14, 4509, 200
×
. Fig. 7. Podobursa triacantha
(Fischli) — ŠJP-4, 1586, 210
×
. Fig. 8. Transhsuum maxwelli (Pessag-
no) — K-2, 4506, 220
×
. Fig. 9. Spongocapsula perampla (Rüst) —
Ps-14, 5842, 175
×
. Fig. 10. Napora lospensis Pessagno — Ps-3,
6999, 190
×
. Fig. 11. Mirifusus dianae (Karrer) — K-1, 4567, 135
×
.
Fig. 12.
Fultacapsa aff. sphaerica (Ožvoldová) nov. comb., the apical
view of Fig. 3 — Ps-14, 3052, 300
×
. Fig. 13. Podocapsa amphi-
treptera
Foreman — Ps-14, 4266, 145
×
. Fig. 14. Hsuum cuestaense
Pessagno — ŠJP-4, 8341, 210
×
. Fig. 15. Archaeodictyomitra cf. api-
arium
(Rust) — ŠJP-4, 8314, 380
×
. Fig. 16. Archaeodictyomitra cf.
minoensis
(Mizutani) — ŠJP-4, 8312, 290
×
.
Plate VI: Fig. 1.
Emiluvia salensis Pessagno — Ps-3, 5821, 145
×
.
Fig. 2.
Paronaella broennimanni Pessagno — Ps-14, 4278, 160
×
.
Fig. 3.
Napora lospensis Pessagno — ŠJP-4, 8317, 155
×
. Fig. 4.
Paronaella pristidentata
Baumgartner — Ps-14, 4274, 150
×
.
Fig. 5.
Haliodictya
(?) antiqua (Rüst) — K-1, 4585, 110
×
. Fig. 6.
Tetratrabs bulbosa
Baumgartner — ŠJ DD-9, 4512, 75
×
. Fig. 7.
Paronaella mulleri
Pessagno — K-1, 4582, 125
×
. Fig. 8. Emiluvia
salensis
Pessagno — ŠJP-4, 8316, 150
×
. Fig. 9. The transitional
form between Triactoma blakei (Pessagno) and Triactoma jonesi
(Pessagno) — K-2, 4589, 180
×
. Fig. 10. Podocapsa amphitreptera
Foreman — ŠJP-4, 8331, 145
×
. Fig. 11. The transitional form be-
tween Triactoma blakei (Pessagno) and Triactoma jonesi (Pessag-
no) — Ps-14, 4272, 130
×
. Fig. 12. Tetratrabs bulbosa Baumgart-
ner — K-1, 4573, 65
×
. Fig. 13. Podobursa spinosa (Ožvoldová)
— ŠJP-4, 1585, 130
×
. Fig. 14. Paronaella sp. — Ps-3, 5820, 145
×
.
Fig. 15.
Triactoma sp. — ŠJP-4, 8318, 160
×
. Fig. 16. Emiluvia
salensis
Pessagno — K-1, 4578, 150
×
. Fig. 17. Tritrabs casmali-
aensis
Pessagno — ŠJ DD-9, 4259, 150
×
. Fig. 18. Acanthocircus
suboblongus
(Yao) — ŠJP-4, 1581, 380
×
.
The important result is the fact that radiolarites with Mn
coatings (Sokolica Radiolarite Formation) extend to the mid-
dle Oxfordian (the presence of the species Podocapsa amphi-
treptera
Foreman in the section ŠJDD).
The association found in the locality Lúčka represents mid-
dle Oxfordian–early Kimmeridgian (UAZones 9–10) (l.c.).
In the locality Kyjov the assemblage in sample K-l repre-
sented a wide stratigraphical range: UAZone 8 – UAZone 10
(middle Callovian–early Kimmeridgian) (l.c.).
The association from the locality Milpoš were very poor.
At the locality Podsadek, near Stará ubovňa, the most sil-
iceous part of the section contain radiolarians indicating a
middle Oxfordian–early Kimmeridgian age (UAZones 9–10)
(l.c.). Distribution of radiolarians in the studied sections
shows Fig. 4.
Systematic part
Genus Fultacapsa Ožvoldová, nov. gen.
Derivatio nominis
: Latin adjective fultus — leant, accord-
ing to the spines, arranged as a support of the test. Latin noun
capsa
— chest. Feminine gender.
Type species
: Acotripus sphericus Ožvoldová l988
Diagnosis
: Nassellariina possesing three or four chambers.
Shape of proximal part of the test subglobose or conical. Cepha-
lis usually bearing apical horn. Terminal segment large, globose
lacking aperture. On its distal part spines occur. Lattice layer on
terminal segment consists of hexagonal to polygonal pore
frames.
Comparison
: Genus Podocyrtis Ehrenberg, Acotripus
Haeckel and Hiscocapsa O’Dogherty have aperture on the
last segment. The spinose species of the genus Sethocapsa
Haeckel and Birkenmajeria Widz have the spines as the or-
namentation of the pore frames. This new genus only pos-
sesses spines on the distal part of the terminal segment.
Included species
:
Cyrtocapsa
sp. sensu De Wever et al. l986
Sethocapsa
(?) sphaerica (Ožvoldová) l988
Sethocapsa
(?) concentrica (Steiger) l992
Sethocapsa tripes
Yang l993
Range
: Middle Oxfordian – early Tithonian up to date known.
Fultacapsa sphaerica
(Ožvoldová l988) nov. comb.,
emend. diagnosis Pl. V: Figs. 1, 2
l988 Acotripus sphericus n.sp. — Ožvoldová, p. 376, pl. 5, Fig. 1–5, 7
l993 Birkenmajeria sphaerica (Ožvoldová) — Widz-De Wever, p. 82,
pl. 1, Fig. 3–4
l995 Sethocapsa (?) sphaerica (Ožvoldová) — Baumgartner et al.,
p. 500, pl. 3168, Fig. 1–4
Emended diagnosis:
The apical horn of variable thickness
and length rises from cephalis. The apical horn and tips on
the terminal segment can be split into three small spines
(Pl. V: Figs. 1, 2 in this article. Pl. 1, Fig. 4 in Widz & De
Wever l993).
Remarks:
In the type species (Ožvoldová l988) the exist-
ence of apical horn was indistinct. But this character was
confirmed in other occurrences of this species, (Danelian
1989, pl. 2, Fig.1–5, Mišík et al. l994, pl. 1, Fig. 16).
archives GÚDŠ
, Bratislava (in Slovak).
Range:
Middle Oxfordian – early Tithonian according to
Baumgartner et al. (1995)
§
60 OŽVOLDOVÁ and FRANTOVÁ
Fig. 4.
Distribution of radiolarians in the studied sections (Ps — Podsadek, ŠJP, ŠJDD — Šarišské Jastrabie, K — Kyjov, L — Lúčka,
M — Milpoš).
SJ
SJ
SJ
SJ
SJ
Ps
Ps SJP SJP SJP DD DD DD DD DD
K
K
L
M
D istr ibu tio n o f r a d io la ria n s in stu d ied sa m p les
3
14
2a
4
1
4
5
6
8
9
1
2
5
5
Acaeniotyle diaphorogona Forem an
*
*
*
*
Acanthocircus suboblongus (Yao)
*
*
*
*
Acanthocircus trizonalis (Rust)
*
Archaeodictyomitra cf. apiarium (Rust)
*
*
*
Archaeodictyomitra directiporata (Rust)
*
Archaeodictyomitra cf. minoensis (M izutani)
*
Archaeodictyomitra prisca Kozur et M ostler
*
*
Archaeospongoprunum imlayi Pessagno
*
*
*
*
*
*
Archaeospongoprunum sp.
*
*
*
Angulobracchia biordinalis Ovoldová
*
Angulobracchia digitata Baumgartner
*
Angulobracchia (?) cf. rugosa Jud
*
Bernoullius dicera (Baumgartner)
*
Cinguloturris carpatica Dum itrica
*
*
Emiluvia chica Foreman
*
Emiluvia orea Baumgartner
*
*
*
*
*
Emiluvia ordinaria Ovoldová
*
*
*
Emiluvia pessagnoi s.l. Foreman
*
*
*
Emiluvia pessagnoi multipora Steiger
*
Emiluvia p+A35remyogii Baum gartner
*
*
*
Emiluvia salensis Pessagno
*
*
*
*
*
*
Emiluvia sedecim porata (Rust)
*
*
*
Eucyrtidiellum ptyctum (Riedel et Sanfilippo)
*
*
Fultacapsa sphaerica (Ovoldová)
*
*
*
*
*
*
Fultacapsa cf. sphaerica (Ovoldová)
*
Fultacapsa aff. sphaerica (Ovoldová)
*
Halesium sp.
*
*
Haliodictya (?) hojnosi Riedel et Sanfilippo
*
Haliodictya (?) antiqua (Rust)
*
*
*
*
Hexastylus sp.
*
Hsuum cuestaense Pessagno
*
M irifusus dianae (Karrer)
*
*
*
*
M irifusus cf. fragilis Baum gartner
*
Napora lospensis Pessagno
*
*
Orbiculiforma (?) catenaria Ovoldová
*
*
Orbiculiforma (?) cf. catenaria Ovoldová
*
Paronaella broennimanni Pessagno
*
*
*
Paronaella mulleri Pessagno
*
*
*
*
Paronaella pristidentata Baumgartner
*
Paronaella pygmaea Baumgartner
*
Parvicingula cf. dhimenaensis Baum gartner
*
*
Parvicingula cf. m ashitaensis M izutani
*
Perispyridium ordinarium (Pessagno)
*
Podobursa spinosa (Ovoldová)
*
*
*
*
*
*
*
*
*
*
Podobursa triacantha (Fischli)
*
*
*
*
*
*
*
Podobursa polyacantha (Fischli)
*
Podocapsa am phitreptera Forem an
*
*
*
Pseudodictyom itra cf. prim itiva M atsuoka et Yao
*
*
Pseudodictyom itrella hexagonata (Heitzer)
*
*
Sethocapsa sp.
*
Spongocapsula perampla (Rust)
*
*
*
Syringocapsa spinellifera Baum gartner
*
Tetraditryma pseudoplena Baum gartner
*
*
Tetratrabs bulbosa Baumgartner
*
*
*
*
Tetratrabs zealis (Ovoldová)
*
*
Transhsuum brevicostatum (Ovoldová)
*
*
*
*
*
Transhsuum m axwelli (Pessagno)
*
*
Triactom a blakei (Pessagno)
*
*
*
*
*
*
*
*
Triactom a jonesi (Pessagno)
*
*
*
*
Tricolocapsa yaoi M atsuoka
*
*
*
Tricolocapsa undulata (Heitzer)
*
Triactom a sp.
*
Tritrabs casm aliaensis (Pessagno)
*
*
Tritrabs exotica (Pessagno)
*
*
*
*
*
Tritrabs rhododactylus Baum gartner
*
*
*
*
*
W illiriedellum sp. A sensu M atsuoka 1983
*
Zham oidellum ovum Dumitrica
*
*
*
JURASSIC RADIOLARITES FROM EASTERN PART OF PIENINY KLIPPEN BELT 61
Fultacapsa aff. sphaerica
(Ožvoldová 1988)
Pl. 5, Fig. 3, 12
Remarks:
The specimen differs from the holotype by the
subglobular instead conical shape of the first three (?) seg-
ments and by the existence of much shorter spines on the ter-
minal segment.
Occurrence:
Ps-l4 (late Oxfordian–early Kimmeridgian —
UAZone 10 sensu Baumgartner et al. 1995).
Acknowledgements:
The authors would like to express their
gratitude to Dr. Luis O’Dogherty (University of Cadiz) for
thoroughly reviewing the manuscript and kindly improving
its language. We would like also thank to Prof. Mišík (Come-
nius University of Bratislava) for his review.
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